Bellardithala laubei ( Hoernes & Auinger, 1880 ) Harzhauser & Landau, 2021

Harzhauser, Mathias & Landau, Bernard, 2021, An overlooked diversity-the Costellariidae (Gastropoda: Neogastropoda) of the Miocene Paratethys Sea, Zootaxa 4982 (1), pp. 1-70 : 20-22

Bellardithala laubei ( Hoernes & Auinger, 1880) nov. comb.

Figs 7F 1 –F 2

[ Callithea ] Laubei nov. form.—Hoernes 1880: 125 [nomen nudum].

* Mitra Laubei nov. form.— Hoernes & Auinger, 1880: 89 (pars), pl. 10, figs 19a–c.

Mitra (Thala) laubei Hö. Au. — Boettger 1906: 11.

Th [ala]. laubei (R. Hörn, et Au.)— Sieber 1958a: 153.

Type material. Lectotype (designated herein): NHMW 1999 z0076/0002, SL: 8.2 mm, MD: 2.4 mm, Bad Vöslau ( Austria), illustrated in Hoernes & Auinger (1880, pl. 10, figs 19a–c), figs 7F 1 –F 2.

Revised description. Shell medium-sized, very slender fusiform. Protoconch unknown. Teleoconch of 5.5 weakly convex whorls with periphery slightly above abapical suture. Suture incised. First teleoconch bearing about 11 opisthocline axial ribs, separated by narrow interspaces, crossed by faint spiral grooves. Spiral grooves strengthening slightly on later whorls, forming spiral rows of blurred nodes at intersections with axial ribs. Spiral cord at adapical suture most prominent, followed below by two spiral rows of nodes. Nodes on subsequent cords indistinct, blurred and largely reduced below periphery. Base slowly contracting. Aperture elongate, narrow, anal sinus indistinct. Narrow columellar callus. Columella with four oblique folds. Outer lip weakly thickened with faint denticles within. Siphonal canal long and wide, bent slightly to left, with shallow, wide siphonal notch.

Shell measurements and ratios. SL: 8.2 mm, MD: 2.4 mm, AA: 29°, SL/MD: 3.5, AL/AW: 6.94, AH/S: 2.3–2.4.

Discussion. Hoernes & Auinger (1880) based Mitra laubei on a series of syntypes, which, in our opinion, are not conspecific. As first revisers we designate specimen NHMW 1999z0076/0007 from Bad Vöslau ( Austria), illustrated by Hoernes & Auinger (1880, pl. 10, figs 19a–19c) as lectotype. All other specimens are described herein as new species ( Bellardithala baluki , B. dacica ). Bellardithala laubei ( Hoernes & Auinger, 1880) differs from all of its Paratethyan congeners in having a far more slender fusiform outline and slightly more convex whorls. In sculpture it is similar to B. cf. dacica nov. sp., but that species is broader fusiform, and the spire whorls are slightly scalate in profile.

Palaeoenvironment. The clay at Bad Vöslau formed in middle neritic settings with up to 250 m water depth ( Hohenegger et al. 2008).

Distribution in Central Paratethys. Badenian (middle Miocene): Vienna Basin: Bad Vöslau ( Austria).

Figures

FIGURE 1. A. Map of central and southeastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005) and Nosowska (2020). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E–Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Measurements to shell morphology and whorl profiles. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height. AL: aperture length. AW: aperture width. S: length of siphonal canal.

FIGURE 7 A1–A2. Bellardithala baluki nov. sp., holotype, Vienna/Pötzleinsdorf (Austria), NHMW 1859/0047/0015. B1–B2. Bellardithala baluki nov. sp., paratype, Vienna/Pötzleinsdorf (Austria), NHMW 1859/0047/0015b. C1–C2. Bellardithala baluki nov. sp., paratype, Vienna/Pötzleinsdorf (Austria), NHMW 2020/0063/0001. D1–D2. Bellardithala fedosovi nov. sp., holotype, Steinebrunn (Austria), NHMW 1864/0001/0563. E1–E2. Bellardithala fedosovi nov. sp., paratype, Steinebrunn (Austria), NHMW 2020/0052/0001. F1–F2. Bellardithala laubei (Hoernes & Auinger, 1880), lectotype, Bad Vöslau (Austria), NHMW 1999z0076/0002. G1–G2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 1866/0001/0806. H1–H2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 2020/0054/0001. I1–I2. Bellardithala cf. dacica nov. sp., Niederleis (Austria), NHMW 2020/0054/0002.