Eustrophus tomentosus Say 1826

Pollock, Darren A., 2012, Review of the Eustrophinae (Coleoptera, Tetratomidae) of America north of Mexico, ZooKeys 188, pp. 1-153 : 12-13

Eustrophus tomentosus Say 1826 Figures 41729415067

Mycetophagus niger Melsheimer 1806: 14 [Catalogue]; Melsheimer 1846: 58.

Mycetophagus tomentosus Melsheimer 1806: 14 [catalogue].-( “Pennsylvania”); Say 1826: 239.

Eustrophus niger Melsheimer 1846: 58.- Melsheimer 1853: 143 (syn.); Leng 1920: 238; LeSage 1991: 246; Poole and Gentili 1996: 299.

Eustrophus tomentosus Say 1826: 239.-U.S.A., Illinois (no specific localities given in description); Melsheimer 1853: 143; Crotch 1873: 112; Provancher 1877: 466; Henshaw 1885: 124; Horn 1888: 35; Blatchley 1910: 1293; Leng 1920: 238; Csiki 1924:10; Hatch 1965: 66; LeSage 1991: 246; Poole and Gentili 1996: 299; Young and Pollock 2002: 416; Majka and Pollock 2006: 53; Pollock 2008: 273, 290; Majka and Pollock 2010: 455.

Diagnosis.

The following combination of characters is diagnostic for this species: widely separated eyes; antennal sensilla interrupted, not completely annular; dorsal setae distinctly golden to brown; meso- and metatibiae with oblique ridges.

Description

(from Pollock 2008: 273). TL 4.5-6.0 mm; GEW 2.1-3.0 mm. Body oval, parallel-sided (Fig. 4), distinctly convex dorsally (Fig. 17); dorsal color dark brown, with golden sheen due to dense pubescence; venter and antennae uniformly dark rufous, lighter than dorsal color; dorsal pubescence relatively short, but dense, giving distinct sheen (almost iridescent); eyes widely separated (space ~1.5 × length of first antennomere), inner margins deeply emarginate (Fig. 29); antennomeres 2-11 only slightly but evenly widened to apex, without distinct change in size between any 2 adjacent antennomeres; distal antennomeres subtriangular to nearly quadrate; antennal sensilla not completely annular, present on short sides of antennomeres only; last maxillary palpomere slightly securiform; prosternal process (Fig. 41) triangular, narrowed distally, extended to slightly short of posterior margin of procoxae; prothoracic episternal suture absent (Fig. 50); elytral punctation relatively fine, punctures arranged in longitudinal striae; meso- and metatibiae with oblique ridges present.

Distribution

(Fig. 67). The distribution of this species is virtually transcontinental, with a gap in the interior; for example, no specimens are known from Saskatch ewan and Alberta. Most records from the United States are eastern, but specimens are also known from the Pacific Northwest, California, and Arizona. The 983 specimens examined in this study are from the following: CANADA: BRITISH COLUMBIA, MANITOBA, NOVA SCOTIA, ONTARIO, QUEBEC. UNITED STATES: ALABAMA: Lee, Madison, Marion, Mobile. ARIZONA: Coconino, Pima. ARKANSAS: Crawford, Faulkner, Fulton, Logan, Polk, Pulaski. CALIFORNIA: Butte, Calaveras, El Dorado, Placer, Yuba. COLORADO. CONNECTICUT: Litchfield. DELAWARE: Sussex. DISTRICT OF COLUMBIA. FLORIDA: Dixie, Monroe. GEORGIA: Bartow, Clarke, Echols. IDAHO: Boundary, Clearwater, Latah. IOWA: Johnson, Keokuk, Story. ILLINOIS: Bond, Champaign, Clark, Coles, Cook, LaSalle, McHenry, St. Clair, Union, Wabash. INDIANA: Montgomery, Parke, Porter, Tippecanoe, Vigo. KANSAS: Douglas. KENTUCKY: Butler, Henderson. LOUISIANA: Natchitoches. MARYLAND: Anne Arundel, Charles, Dorchester, Montgomery, Prince George’s, Talbot. MASSACHUSETTS. MICHIGAN: Berrien, Charlevoix, Fillmore, Gogebic, Ingham, Kalamazoo, Kent, Lapeer, Leelanau, Livingston, Macomb, Midland, Monroe, Oakland, St. Joseph, Shiawassee, Washtenaw, Wayne. MINNESOTA: Crow Wing, Hennepin, Sherburne, Washington. MISSISSIPPI: George, Pearl River. MISSOURI: Boone, Dent, Franklin, Randolph, St. Charles. MONTANA: Dawson, Lake, Lincoln, Missoula, Ravalli, Rosebud, Sanders. NEBRASKA: Douglas, Sarpy. NEW HAMPSHIRE: Carroll. NEW JERSEY: Burlington, Gloucester. NEW MEXICO: San Miguel. NEW YORK: Erie, Essex, Genesee, Niagara, Onondaga, Orleans, St. Lawrence, Tompkins. NORTH CAROLINA: Durham, Moore, Stokes, Wake. NORTH DAKOTA: Pembina, Richland. OHIO: Athens, Butler, Clinton, Delaware, Hocking, Ottawa, Portage, Scioto, Vinton, Wayne, Wyandot. OKLAHOMA: Latimer, Oklahoma. OREGON: Benton, Crook, Douglas, Harney, Jackson, Lane, Linn, Multnomah, Wallowa, Yamhill. PENNSYLVANIA: Allegheny, Tioga, Westmoreland. SOUTH CAROLINA: Anderson, Florence, Pickens. TENNESSEE: Shelby. TEXAS: Angelina, Brewster, Montgomery, Sabine, Walker. UTAH: Washington. VERMONT: Franklin. VIRGINIA: Alexandria, Hampton, Nelson, Shenandoah. WASHINGTON: Walla Walla. WEST VIRGINIA: Braxton, Doddridge, Greenbrier, Hampshire, Jackson, Kanawha, Lincoln, Mason, Mineral, Nicholas, Pendleton, Randolph, Ritchie, Roane, Taylor, Webster, Wirt, Wood. WISCONSIN: Dane, Racine, Sauk, Washington. (Complete label data given in Appendix 1).

Types.

Eustrophus tomentosus Say. NEOTYPE (designated by Pollock 2008), sex unknown, labeled "Ill. / E. tomentosus Say. niger Mels.", in MCZ (LeConte collection).

Natural history.

Label data: Pseudotsuga taxifolia (BC), from fungus (BC), evening flight, 4.ix (BC), ex Populus trichocarpa (BC), in bark Larix occidentalis (BC), fleshy fungus on tree (ON), under bark of pine (ON), under wet moldy bark on dead tree (ON), elm (QC), uv light trap (QC), cut surface of stump (QC), Ulmus americana (QC), under bark of pine (AL), Spongipellis unicolor (AR), Climacodes septentrionale (AR), Trametes versicolor (AR), under pine bark (CA), Lindgren trap with turpentine bait (CA), on Pinus ponderosa (CA), ex. Polyporus fungus (FL), uv light in oak-maple forest (IA), fungus on dead pine (ID), under bark of old pine stump (MD), in Peromyscus nest debris under bark of dead standing Liriodendron (MD), fungus on oak (MN), funnel traps (MT), at black light, vi.1970 (NC), malaise trap, viii-ix (NE), window trap, 8-14.vi (NH), light trap, vi (NY), Lindgren funnel trap, alpha-pinene and ethanol lure (OR), stale molasses trap (SC), at wound on oak trunk (SC), malaise in mature hardwood forest, vi (SC), under bark of old dead decid. tree (VT), ex dead oak stump (WI), under bark of chestnut oak (WV). According to Chantal (1985), Eustrophus tomentosus are found under bark of dead trees, especially Ulmus americanus , as well as at sap exudations.