Dothenia, 2002

Sime, Karen R. & Wahl, David B., 2002, The cladistics and biology of the Callajoppa genus-group (Hymenoptera: Ichneumonidae, Ichneumoninae), Zoological Journal of the Linnean Society 134 (1), pp. 1-56 : 24

DOTHENIA WAHL & SIME , GEN. N.

Type-species. Dothenia hansoni sp. n.

Description. Flagellum of female lanceolate, with flagellomere 3 1.3–1.8¥ as long as wide; flagellum of male lacking tyloids. Clypeus flat and wide, apicolateral margin forming an angle of approximately 90°, punctures evenly distributed, apical 0.3 thinned, apical margin straight and with weak median projection. Mandible long and evenly tapered to apex, ventral tooth in same plane as dorsal tooth. Supra-antennal area without paired median ridges or denticles. Vertex with posterior section about as long as ocellar triangle, gently sloping to occipital carina. Gena swollen in dorsal view. Juncture of hypostomal and occipital carinae separated from mandible by about 0.5¥ basal mandibular width. Occipital carina with ventral region above mandible not produced as a low flange.

Pronotal flange low and with posterior face concave. Epicnemial carina dorsally complete. Scutellum strongly convex, often shelf-like, or strongly convex with planar surfaces and forming a more or less acute angle in lateral view; lateral carinae absent. Central convexity of metanotum not widened and lateral depressions not reduced. Base of propodeum rising steeply and abruptly. Areola reduced and partially to completely filled-in, latter state appearing as polished boss. Anterior transverse carina complete, median section not swollen; posterior transverse carina complete; lateral longitudinal carinae complete; median longitudinal carinae, basad anterior transverse carina, either close together on raised areas or fused into single carina; median longitudinal carinae, apicad anterior transverse carina, present or absent. First lateral area with regularly distributed punctures and second lateral area rugosopunctate, or both finely and contiguously punctate. Metapleuron weakly rugosopunctate. Juxtacoxal carina present or absent. Propodeal lateral profile steeply sloping or (in one species) evenly convex. Scopa absent. Cell 1 + 2Rs of fore wing petiolate, vein 2/Rs <3r-m, vein 2 m-cu interception either at midpoint or apicad midpoint of posterior margin (vein 2/M 1.5–2.3¥ as long as vein 3/M).

MS 1 in lateral view with petiole flattened, postpetiole convex; sclerotized anterior portion of S1 0.9 –1.4¥ as long as distance from posterior margin of anterior portion to apex of T1 ; petiole basally without dorsal or lateral bulges; postpetiole with or without basal convexity, median longitudinal carinae absent, median field present, punctures fine and dense and without rugae. Ventral margin of T2 simple. T2-4 without longitudinal rugae. T2-4 evenly convex, with fine contiguous punctures. T2-5 not basally constricted and without dorsolateral ridges. S2 divided and S3-5 entire, or S2-5 entire. Apex of female metasoma with T7-8 barely projecting beyond apex of T6 ; amblypygous

Autapomorphies. Propodeum with median longitudinal carinae anteriad apical transverse carina close together on raised area (#32–1); metapleuron with small contiguous/confluent punctures and without rugae (#36–2); cell 1 + 2Rs of fore wing with vein 2 m-cu interception apicad midpoint of posterior margin (#41–1).

Etymology. The generic name is a euphonious combination of letters with no meaning. Its gender is feminine.

Species and distribution. There is one described species, D. hansoni , and nine undescribed species [AEIC, CCCP, IMLA, JHIC, NHML, UCDC], distributed from Mexico to Argentina.

Biology. The holotype and paratype specimens of D. hansoni (see below) were reared as larval-pupal parasitoids of the sphingid Erinnyis ello (L.). The hosts were collected as final-instar larvae on Euphorbiaceae (D. Janzen, pers. comm.). In addition, this parasitoid has been reared from Manduca occulta (Rothschild & Jordan) on Cestrum glanduliferum (Solanaceae) ( Costa Rica: Area de Conservación Guanacaste, “91- SRNP-1260” [JHIC]), also collected in the final instar (D. Janzen, pers. comm.).