Daealus tuberosus, Zimmerman, 1942

Zimmerman, Elwood C., 1942, Curculionidae of Guam, Insects of Guam I, Honolulu, Hawaii: Bernice P. Bishop Museum, pp. 73-146 : 111-113

23. Daealus tuberosus , new species (pl. 2,D).

Derm reddish brown to black, usually quite shiny where exposed, densely squamose above and below, scaling entirely or almost white or grayish whitein in old faded specimens but well colored in fresh examples as follows: head pale brown, prothorax basically yellowish white with a brownish-yellow cloud at basal halfat at sidesof of disk and one<111 either side of median line; elytra with a broad sub-V-shaped fasciaof of yellowish white scales from sides near or distant from base to suture at about middle, pale scaling continued as a vitta down first two intervals to apex, otherwise mostly with brownish yellow scales but with a dark brown or black elongate patch on third intervalin in frontof TODO and behind sub-V-shaped macula, but these dark patches subject to much variation, sometimes front ones wanting and occasionally with more dark scaling toward apexof of third interval; scaling on legs and underside rather uniformly yellowish white or brownish yellow.

Head with crown densely squamose, derm mostly concealed by scaling, scales erect or suberect, with numerous, spatulate, erect setae scattered throughout that project slightly above scales; front with similar vestiture to crown, with a distinct elongate interocular fovea that usually begins above a line drawn between tops of eyes and terminates near middle of eyes where distinct median carina of rostrum begins, interocular area fully as broad as base of rostrum. Rostrnin with ventral edge three fourths as long as pronotum in male, more than three fourths as long in female, antennae inserted slightly beyond apical third in male, at or slightly behind apical third in female, coarsely and deeply sculptured from base almost to antennae, with three coarse, somewhat irregular dorsal carinae, striae between them densely set with erect setae, coarsely sculptured and carinate only within basal half in female. Aiitennae with scape somewhat longer than funicular segments 1-7 plus basal segment of club; funicle with two basal segments equal in length or first slightly longer, 2 slightly shorter than 3 plus 4, 3 almost as long as 4 plus half of 5; 4 slightly longer than 5, 5 to 7 each successively slightly broader; club as long as segments 4 to 7 plus part of three. Prothorax more than three fourths as long as broad, broadest somewhat beyond middle, base subtruncate, strongly convex on sides from base to apical third or beyond and there with a distinct, not very deep, but variable, subapical constriction, the constriction continued broadly across dorsum which is strongly convex; disk with median line rather deeply and conspicuously impressed and without tubercles, with numerous, scattered, rounded, prominent, polished tubercles extending through scaling; scales similar to those on elytra, very dense, concealing derm, with numerous erect, spatulate setae scattered throughout. Elytra four fifths or five sixths as broad as long, base subtruncate, broadly arcuate on sides from base to about middle and thence rather rapidly narnowed to the apex, longitudinal dorsal contour convex, reaching a summit at about middle and there hardly higher than highest part of pronotum; striae mostly concealed by scaling, much narrower than intervals, their punctures well separated, very small behind basal third and each bearing a decumbent squamiform seta; intervals prominently convex, some of them A-shaped in cross section, especially 3, 5, and 7, interval 1 with a single row of stout, erect, spatulate setae, sometimes setae beginning at a distance from base, without tubercles, 2 with a complete row of prominent tubercles usually interspersed with spatulate setae, 3 usually distinctly more elevated than 2 from base to behind middle, sometimes more strongly elevated at base and just in front of middle, more elevated areas with denser, erect, spatulate setae giving a fasciculate appearance, either entirely without tubercles or with few to many at base, 4 tuberculate, 5 setose, either not tuberculate or with a variable number of tubercles at base, 6-10 tuberculate, tubercles becoming smaller on outer intervals. Legs with femora and tibiae densely squamose and setose; tibiae not carinate. Stertium densely squamose throughout; mesosternal receptacle obviously open in male, its margins fringed with long hair, hind wall slightly more protuberant in female, narrowly and just perceptibly cavernous and without hair; metasternum only one fourth or less as long between mid and hind coxae as breadth of a mesocoxa, squamose only in female, but squainose and with dense long hair in male. Venter with first ventrite tumid and squamose in female, depressed and with disk free from scales and there with long, dense, simple or compound erect hair in male; ventrites 2 to 5 squamose in female, 2 with some long hair and 5 with depressed disk free from scales and hirsute in male. Length, 4-8.0 mm.; breadth, 2-4.0 mm.

Holotype male, Agat, May 31, 1936, Usinger ; allotype female, Machanao, June 5, Usinger ; and the following paratypes: one, same data as holotype; one same place and date, from Hernandia, Swezey ; one, Tarague, May 17, Swezey ; one, Agana , May 25, Usinger ; two, Machanao , from Piper , June 4, Swezey ; one, Machanao , June 5, Usinger ; one, Sinajana , June 15, Swezey ; one, Atao Beach , June 25, Usinger ; two, Machanao , June 30, Swezey ; three, Orote Peninsula, from Pipturus, Sept. 27, Swezey ; and one taken by Fullaway . The following paratypes are in the National Museum material: six specimens taken by Fullaway and labeled only " Island Guam ", one of them taken from Hernandia .

This species varies greatly in size, color, color pattern, and arrangement and number of the tubercles. Old specimens with their bleached scaling appear at first sight to belong to different species than fresh examples. In spite of this variability, specimens of this species have a distinctive facies that is not likely to be confused.