Divellepidosis indubitata ( Skuse, 1888 ) Skuse, 1888

Kolesik, Peter & Gagné, Raymond J., 2016, Revision of early taxa of Australian gall midges (Diptera: Cecidomyiidae), Zootaxa 4205 (4), pp. 301-338 : 318

Divellepidosis indubitata ( Skuse, 1888) , new combination

[ Figs 9 i–n]

Cecidomyia (Colpodia) indubitata Skuse, 1888: 114 .

Colpodia indubitata ( Skuse 1888) , new combination by Gagné (1989b: 155).

Material studied. Holotype male (ANIC 29-38489), “Middle Harbour (Skuse). September”. The type retains the wings, head with partially shrivelled flagellomeres of one complete antenna, thorax with one hindleg, and the abdomen.

Description. Wing 1.8 mm long, 0.5 mm wide; R4+5 strongly curved at apical third, joining C at wing apex, C broken at juncture with R4+5, Rs fully developed, halfway between arculus and end of R1; Cu forked, Cu2 stronger than Cu1; wingfold weak. Head: palpus 4-segmented, palpiger present; flagellomeres 14, with single node, bearing single horizontal circumfilar band, neck 1.5x longer than node. First tarsomere with small blunt apicoventral projection, tarsal claws simple, empodium shrivelled on the remaining leg of holotype. Terminalia: gonocoxites short, without apical lobe dorsally; gonocoxal apodemes separate, ending anteriad of gonocoxites; gonostylus wide basally, with inner edge straight, outer edge curved, abruptly tapering towards apical comb-like tooth, entirely microtrichose and setose; cerci large, bluntly triangular, shallowly divided, with several apical setae; hypoproct concave, slightly shorter than cerci, with single seta on either lobe; tegmen stout, slightly concave apically, longer than cerci, bearing no papillae, ejaculatory apodeme starting between base of gonocoxal apodemes and base of aedeagus and reaching tip of aedeagus.

Remarks. This species and D. pallidina represent the first records of Divellepidosis Fedotova & Sidorenko ( Porricondylinae : Porricondylini ) in Australia. The two species differ in the following characters: in D. pallidina the wing vein Rs is closer to the distal end of R1 than to the arculus, the gonocoxite is long with an acute dorsoapical lobe and the gonoxocal apodemes reach the base of gonocoxites; in D. indubitata R S lies halfway between R1 and the arculus, the gonocoxite has no apical lobe, and the gonoxocal apodemes reach far anteriad of the base of the gonocoxites.

Figures

FIGURE 9. Divellepidosis pallidina (Skuse) (a – h), Divellepidosis indubitata (Skuse) (i – n). (a – f, i – n) male, (g – h) female. (a) terminalia in dorsal view (left cercus lobe in lateral view), (b) wing, (c) sixth flagellomere in ventral view, (d) sixth flagellomere in dorsal view, (e) first tarsomere of hindleg, (f) last tarsomere of hindleg with claw and empodium, (g) sixth flagellomere in ventral view, (h) sixth flagellomere in dorsal view, (i) terminalia in dorsal view, (j) sixth flagellomere in ventral view, (k) sixth flagellomere in dorsal view, (l) wing, (m) first tarsomere of midleg, (n) palpus with palpiger.