Acanthocephalonotum martinsnetoi, Petrulevicius, Julian F. & Popov, Yuri A., 2014

Petrulevicius, Julian F. & Popov, Yuri A., 2014, First fossil record of Discocephalinae (Insecta, Pentatomidae): a new genus from the middle Eocene of Rio Pichileufu, Patagonia, Argentina, ZooKeys 422, pp. 23-33 : 24-26

Taxon classification Animalia Hemiptera Pentatomidae

Acanthocephalonotum martinsnetoi sp. n. Figs 1, 2

Diagnosis.

Same as for the genus, by monotypy.

Description.

The specimen is mainly complete and articulated in dorsal position with a composite view of dorsal and ventral structures.

Body: 4.7 mm long and 3.6 mm wide at pronotum; width (at the base of the hemelytra) / specimen length ratio, 0.78; antennae and legs not visible; head broad, almost rectangular with numerous punctures, wider than long; anterior margin of head almost straight in almost all its width; head 1.15 mm wide in its anterior margin, 0.8 mm long; eyes, 0.24 mm wide, 0.11 mm long; anteocular length 0.36 mm; inter-ocular width 0.95 mm; inter-ocular width / head length ratio, 0.84; distance between ocelli 0.48 mm; distance between eyes and ocellus 0.2 mm; juga (= mandibular plates) touching each other before clypeus; apex of juga contiguous about 0.11 mm; lateral margins of juga deeply concave; clypeus bullet shaped; ante-ocular process extending beyond the eye and perpendicular to the sagittal plane, subtriangular shaped, 0.23 mm long, with its anterior margin convex and posterior margin concave and beside the eye; labrum thick and curved (ventral structure); origin of the labium caudad of the anterior limit of the eyes (ventral structure); pronotum with a broad spine-like antero-lateral process, stout and acute, parallel to the sagittal plane, 0.2 mm long; head length / pronotal width ratio, 0.87; pronotum with numerous punctures, strongly explanate and bean-shaped, 3.6 mm wide, 1 mm long; lateral margins rounded and irregular; scutellum triangular with a developed and circular tongue; scutellum about 2.8 mm wide at base, 1.9 mm long; tongue, 1.2 mm wide and 0.75 mm long; apex of tongue surpassing the corium; apex of scutellum reaching the anterior side of abdominal segment 7; posterior margin of abdominal segment 7 with three straight sides; gonocoxites 8 (ventral structure) with sub-triangular truncate shape, outer lateral margins obliques, posterior ones straights; laterotergites 8 large, sub-triangular, truncate in inner lateral margins.

Wings: well developed membrane just surpassing end of abdomen; corium with punctures; costal margin bending acutely before end of basal half (boomerang shaped); costal angles of corium above ante-penultimate tergum; R slightly curved and followed by punctures by both sides; M slightly zigzagged; CuA almost straight and followed by punctures by both sides; venation not visible in membrane.

Material.

holotype specimen MAPBAR 4137.

Type locality and horizon.

Volcanic caldera-lake beds, Río Pichileufú, quarry RP3 ( Wilf et al. 2005), province of Río Negro, Patagonia Argentina, palaeolatitude ~ 46°S; middle Eocene (47.7 Ma) ( Wilf et al. 2005; Wilf 2012).

Etymology.

Dedicated to the memory of Rafael Gioia Martins-Neto, outstanding palaeoentomologist and “irmão de alma", who unexpectedly and prematurely passed away in 2010 at age 56.