Campsicnemus leucostoma Evenhuis

Evenhuis, Neal L., 2012, Review of the Campsicnemus fumipennis group (Diptera: Dolichopodidae) in the Hawaiian Islands, with descriptions of new species and corrections of misidentifications, Zootaxa 3497, pp. 1-16 : 9-11

Campsicnemus leucostoma Evenhuis , sp. nov.

( Figs. 4 , 6 , 9 , 14 , 18 )

Diagnosis. Similar to C. aniani and can be separated from it by the all generally white body except the occipital region of the head, the admedian vittae and brown tergal pattern (these white areas being yellowish brown to dark brown in C. aniani ).

Description. Male: Body length: 3.0–3.5 mm. Wing length: 3.2–3.8 mm. Head: Vertex, occiput and postgena brown, remainder of head white; face and clypeus silvery tomentose; oc and vt brown, about two-thirds length of antennal arista; postgena with fine white hairs; face constricted at middle, holoptic for distance of 2 ommatidia; palpus small, white; proboscis white, extending below eye in lateral view; antenna ( Fig. 4 ) with segments white; postpedicel subtriangular, length about 1.5X width, rounded apically; arista slightly longer than head height. Thorax: White except as follows: anepimeron brown at wing base; laterotergite with small brown spot laterally; pair of admedian vittae brown, prescutellar and prealar areas diffuse pale brown; scutellum brown dorsally; thoracic setae very long, brown: 1 + 4 dc; 2 np; 2 ph; 1 pa; 1 + 1 + 1 sc (strong scutellar as long as head height); 10–12 ac; halter stem and knob white. Legs: White, coxae and femora with white hairs except ventral femoral hairs brown; FII with 20–25 setae ventrally, yellow, longest and thinnest on basal third, stiff, brown and shorter on apical two-thirds; TiII ( Fig. 18 ) long, straight, with 4 long stiff, black setae on lateral surface, sparse patch of long hairs subbasally on lateral surface (MSSC), stiff hairs on medial surface, shortest on basal and apical third, longest on medial third, hairs erect on basal two-thirds, becoming appressed on apical third; apical tibial setae not as strong as in C. aniani or C. fumipennis . IIt1 about 1.5X length of IIt2, without MSSC. Remaining leg segments without MSSC. Wing ( Fig. 14 ): Hyaline; posterior crossvein almost 1/3 length of distal section of vein CuA1. Abdomen: Brown dorsally, with white laterally; short brown hairs dorsally on each tergite, hairs yellowish brown laterally, hairs longest on posterior margin of tergite I; sternites white. Hypopygium yellow with white cerci, not dissected. Female: As in male except for lack of MSSC and as follows: eyes dichoptic, separated by width of one ommatidium; antennal postpedicel length subequal to width; brown areas of thorax and abdomen slightly darker and more extensive than in male.

Types. HOLOTYPE 3 (BPBM 16,756; preserved in spirit) and 13, Ƥ paratypes from HAWAIIAN ISLANDS: Hawai‘i: Pu‘u Maka‘ala Natural Area Reserve, 1330 m, 4–10 Apr 1994, Malaise trap, D.J. Preston.

Other paratypes: HAWAIIAN ISLANDS: Hawai‘i: 1Ƥ, Pu‘u Maka‘ala Natural Area Reserve, Army Road 4, 1 mi S Stainback Hwy, 19°33.149'N, 155°13.849'W, 11–12 Aug 2012, yellow pan trap in ‘ ō hi‘a/Cybotium forest, N. Evenhuis (BPBM); 13, Upper Ola‘a Forest Reserve, 4000 ft [1020 m], 16 Apr 1991, reared ex Cheirodendron leaves, K.Y. Kaneshiro (UHM); 13, Kona Hema TNC Nature Preserve, Papa Unit, 19°12'47.217"N, 155°49'41.625"W, 11 Jan 2006, Malaise trap MT1, R. Peck (BPBM); 13, Hawaii Volcanoes National Park, Ola‘a Forest, Pole 44, 29 Jul 2010, general sweeping, B. Ort (BPBM), 43, 15Ƥ, Hawaii Volcanoes National Park, Ola‘a forest, SEA, Pu‘u Unit, 4200 ft [1280 m], Jun–Aug 1993, 3185-001, 3929-002, 2333-001, 3845-002, 2111-001, 2636-003, 2829-004, 2352-001, 2341-006, 2327-003, 3021-002, 2316-001, 2249-001, green pan traps, D.A. Foote, N.L. Evenhuis (HAVO).

Etymology. The specific epithet derives from the Greek leucos = “white” + stoma = “mouthparts”; referring to the distinctive white mouthparts of the species.

Figures

FIGURE 6. Campsicnemus leucostoma, sp. nov., male habitus. Body length: 3.2 mm.

FIGURES 2 – 5. Campsicnemus male antennae. 2, C. aniani, sp. nov.; 3, C. fumipennis; 4, C. leucostoma, sp. nov.; 5, C. spectabulus, sp. nov.

FIGURES 7 – 10. Campsicnemus female thorax and abdomens. 7, C. aniani, sp. nov.; 8, C. fumipennis; 9, C. leucostoma, sp. nov.; 10, C. spectabulus, sp. nov.

FIGURES 12 – 15. Campsicnemus wings. 12, C. aniani, sp. nov.; 13, C. fumipennis; 14, C. leucostoma, sp. nov.; 15, C. spectabulus, sp. nov.

FIGURES 16 – 19. Campsicnemus male mid tibiae. 16, C. aniani, sp. nov.; 17, C. fumipennis; 18, C. leucostoma, sp. nov.; 19, C. spectabulus, sp. nov.