Andrena gomerensis WARNCKE , 1993

Kratochwil, Anselm & I, Canary, 2020, Revision of the Andrena wollastoni group (Hymenoptera, Anthophila, Andrenidae) from the Madeira Archipelago and the Canary Islands: upgrading of three former subspecies and a description of three new subspecies, Linzer biologische Beiträge 52 (1), pp. 161-244 : 190-195

publication ID	https://doi.org/ 10.5281/zenodo.5273217
persistent identifier	https://treatment.plazi.org/id/D91C87FC-FFFB-B14E-FF2A-8EB5F112E3F8
treatment provided by	Marcus
scientific name	Andrena gomerensis WARNCKE , 1993
status

Treatment

Andrena gomerensis WARNCKE, 1993 View in CoL

Syn. Andrena wollastoni gomerensis WARNCKE, 1993 p.p.

S t a t u s: WARNCKE (1993) defined 19 types (10♀♀, 9♂♂). In the OLML, the holotype and seven paratypes (3♀♀, 4♂♂; BLANK & KRAUS 1994) are deposited, and in the MCNSC one paratype (1♀, ORTEGA 2005) is deposited. WARNCKE (1993) described A. w. gomerensis for La Gomera and El Hierro. The paratypes of the OLML are characterised in detail below, the female paratype of the MCNSC is labelled (1♀ Montaña de la Zarza, 1000 m, 20 km WNW S. Sebastián, 03.06.1983, leg. Hohmann; MCNSC, HY-7031). It was not possible to analyse the following paratypes of A. w. gomerensis, because their deposition is unknown: 4♀♀, 2♂♂ Hermigua El Moralito, 17.03.1990; 1♀, 3♂♂ Vallehermoso, Altos de Argamul, 13.03.1990. All the types from La Gomera have been used for the description of A. g. gomerensis.

Initially, WARNCKE (1968) grouped the specimens from La Gomera to A. w. acuta WARNCKE, 1968 . He explained that in WARNCKE (1968) only males were analysed, and, after studying females and further males, the author concluded that the specimens of La Gomera are not A. w. acuta but belong to an own subspecies: A. w. gomerensis. Ten females collected by J. A. W. Lucas from La Gomera in 1992 were labelled by Warncke as A. wollastoni acuta War. (printed), but this was not corrected in A. w. gomerensis. These ten specimens (OLML6-15) are also deposited in the OLML and are now used for the description of A. g. gomerensis.

The name A. w. gomerensis was first mentioned in the unpublished thesis of LA ROCHE BRIER (1992), but A. w. gomerensis WARNCKE, 1992 , is a nomen nudum (the description is missing in the publication). The names cited in this thesis were not available at that date according to Articles 8 and 9 ( ICZN 1999); see BLANK & KRAUS (1994).

HOHMANN et al. (1993) did not mention that one specimen of A. g. gomerensis was also found in El Hierro (paratype: 1♀ El Hierro , Costa de Val Verde, 28.04.1942, collector unknown, ID-No OLML39 ) ; checked by A. Kratochwil. Apart from this detection, A. gomerensis was never found there again despite intensive personal searching, so this specimen might have been wrongly labelled. Therefore, I do not list A. g. gomerensis for El Hierro.

F o r m e r d e s c r i p t i o n s: WARNCKE (1993): First tergite lateral rounded (in A. w. acuta flattened and slightly carinate) and considerably shagreened. In males the first tergite is also rounded, but this is not so apparent by the lesser lateral tergite width. This characterisation was adopted by GUSENLEITNER & SCHWARZ (2002).

D i a g n o s t i c q u a l i t a t i v e f e a t u r e s: Inadditiontothemorphological characteristics that are typical for the taxa of the A. wollastoni group, A. gomerensis is characterised by the following specific features:

F e m a l e: Colour. Head: flagellum black or dark brown ( Fig. 14d View Fig ). Mesosoma: femur, tibia, and basitarsus black (dark brown) (A. g. palmae nov. ssp. partly reddish in 10%) ( Fig. 14a View Fig ); mediotarsi reddish-brown ( Figs. 14a, d View Fig ); wings brownish toned, veins reddish-brown ( Fig. 14f View Fig ); pterostigma dark yellowish or reddish-brown ( Figs. 14a, f View Fig ). Metasoma: T1-4 black with black to dark reddish-brown or dark brown depression zone; T5 depression zone reddish-brown.

Pubescence. Head: clypeus and supraclypeal area with yellowish-white not dense hairs ( Figs. 14c, 14d View Fig , 15b View Fig ); paraocular area with yellowish-white hairs, some brownish (black) hairs between the subantennal socket and the facial fovea; scapus and antennal socket with dorsal longer brownish hairs and ventral shorter yellowish hairs; genal area with white hairs; facial fovea in the upper part with brownish hairs, in the lower part with whiteyellowish hairs ( Fig. 15a View Fig ); vertex behind the ocelli with some longer yellowish-white and brownish hairs. Mesosoma: mesoscutum and scutellum with yellowish-brownish hairs, in some cases laterally with few yellowish hairs; mesepisternum with yellowish-white hairs; propodeal corbicula with some yellowish-white hairs and some hairs in the centre; trochanteral and femoral flocculus with yellowish-white hairs; tibial scopa dorsally with brownish hairs (A. g. palmae nov. ssp. with dorsoventral slightly brownish hairs), dorsobasally with reddish-brown hairs, tibial scopa ventral with yellowish-white hairs ( Fig. 15f View Fig ). Metasoma: tergites scarcely hairy, T2 and T3 (T4) with laterally yellowishwhite, fragmentary open hair bands (dense rows of hairs) or with fragmentary rows of hairs between the tergite and the tergite depression; T4 with row of hairs between tergite and tergite depression; T5 laterally white or yellowish-white hairs, in the centre reddishbrownish hairs, reaching to the pygidium ( Fig. 14f View Fig ), T6: dark brown hairs.

Structure. Head: clypeus convex without (A. g. gomerensis) or with fragmented impunctate line (A. g. palmae nov.ssp. 71%, A. g. gomerensis 29%) ( Fig. 15b View Fig ), clearly shagreened, slightly dull; clypeus with shallow punctures, more or less dense (PD: 28 μm, PDI: 14-56[70] μm) ( Fig. 15b View Fig ); labrum process liguliform-trapezoidal or trapezoidal (A. g. gomerensis) ( Fig. 15b View Fig ) or triangular (A. g. palmae nov.ssp.), rounded on the top. Mesosoma: scattered punctured with very shallow punctures, especially in front (PD: 14- 42 μm) ( Fig. 15c View Fig ); propodeum rugose, primarily in the basal centre and in the dorsolateral area ( Fig. 15d View Fig ). Metasoma: tergites hammertone-like shagreened and shiny; shallow and very scattered but not clearly punctured (PD: 14 μm) ( Fig. 15e View Fig ); T1 no depression zone or only with a slight depression zone, T2-T4 with deep depression zones.

M a l e: similar to female with following differences: Colour. Head: distal half of the mandible not or partly reddened; flagellum brown or (light) brown ( Figs. 16c, d View Fig ). Mesosoma: mediotarsi reddish-brown or brown (A. g. gomerensis) or reddish-brown (A. g. palmae nov.ssp.); wings light brownish toned ( Fig. 16f View Fig ); pterostigma yellowish in the centre, reddish-brown marginated.

Pubescence. Head: clypeus yellowish-white with darker hairs at the base ( Fig. 17b View Fig ); paraocular area with yellowish-white hairs, many brownish (black) hairs near subantennal socket; genal area upper part with brownish hairs, lower part with yellowish-white hairs ( Fig. 16c, d View Fig ). Mesosoma: mesepisternum with long yellowish-white hairs ( Fig. 16c View Fig ).

Metasoma: base of T1 with some yellowish-white hairs, scarcely hairy in the centre, T2- T4 depression zones with laterally yellowish-white, strongly fragmented hair bands (dense rows of hairs); T5, T6 with yellowish or yellowish-reddish hairs; ST 8: with long yellowish hairs at the end.

Structure. Head: vertex above ocelli narrow, as wide as ocellar diameter or half of the diameter ( Fig. 16d View Fig ); clypeus in front deeper, in the centre and base shallow and not densely punctured (PD: 28 μm, PDI: 14-28 μm) ( Fig. 17b View Fig ); labrum process trapezoidal, emarginated, ends left and right side (slightly) thickened ( Fig. 17b View Fig ). Mesosoma: very scattered punctured (PD = 14 μm) ( Fig. 17c View Fig ). Metasoma: T1 slightly carinate, tergites very scattered punctured (PD = 14μm).

S u b s p e c i e s d i f f e r e n t i a t i o n: A. gomerensis is differentiated into two subspecies: A. g. gomerensis WARNCKE, 1993 (La Gomera), and A. g. palmae nov.ssp. (La Palma). In females, morphological differences exist in the labrum process structure, the mesosoma puncture diameter, partly in the existence of a clypeus impunctate line, partly in the colour of the mediotarsi, and partly in the colour of the tergite depression zone. In males, morphological differences exist in the colouration of the flagellum, in the colour of the pubescence of ST8, partly in the diameter of the vertex width, partly in the colour of the the mediotarsi, and especially in morphometric parameters.