Tachyusa obsoleta Casey, 1906

Pa ṡnik, Grzegorz, 2006, A revision of the World species of the genus Tachyusa Erichson, 1837 (Coleoptera, Staphylinidae: Aleocharinae), Zootaxa 1146, pp. 1-152 : 130-134

Tachyusa obsoleta Casey, 1906 ( Figs. 135–137)

Tachyusa obsoleta Casey, 1906: 208 .

Tachyusa dakotana Casey, 1906: 210 , syn. n.

Tachyusa illini Casey, 1906: 210 , syn. n.

Tachyusa missouriana Casey, 1906: 209 , syn. n.

Tachyusa parviceps Casey, 1906: 208 , syn. n.

Tachyusa pruinosa Casey, 1906: 210 , syn. n.

Tachyusa subaluatacea Casey, 1906: 209 , syn. n.

Tachyusa (s. str.) obsoleta: Bernhauer & Scheerpeltz, 1926: 584 (as synonym of T.

cavicollis).

Tachyusa (s. str.) dakotana: Bernhauer & Scheerpeltz, 1926: 586 (as synonym of T. pruinosa ) Tachyusa (s. str.) illini: Bernhauer & Scheerpeltz, 1926: 586 (as synonym of T. pruinosa )

Tachyusa (s. str.) missouriana: Bernhauer & Scheerpeltz, 1926: 584 (as synonym of T.

cavicollis).

Tachyusa (s. str.) parviceps: Bernhauer & Scheerpeltz, 1926: 584 (as synonym of T.

cavicollis).

Tachyusa (s. str.) pruinosa: Bernhauer & Scheerpeltz, 1926: 586 (as valid species).

Tachyusa (s. str.) subaluatacea: Bernhauer & Scheerpeltz, 1926: 584 (as synonym of T.

cavicollis).

Type material. Tachyusa obsoleta Casey. Holotype: ♂: North Carolina, Asheville , type USNM 38948 ( USNM).

Tachyusa dakotana Casey. Holotype: ♂: North Dakota, Williston, June 8–9, Wickham , type USNM 38952 ( USNM).

Tachyusa illini Casey. Holotype: ♀: Missouri, Creve Coeur Lake , type USNM 38954 ( USNM).

Tachyusa missouriana Casey. Holotype: ♀: Missouri, type USNM 38953 ( USNM).

Tachyusa parviceps Casey. Holotype: ♀: Pennsylvania, type USNM 38949 ( USNM).

Tachyusa pruinosa Casey. Lectotype (here designated): ♀: New York, Catskill Mts. , type USNM 38951 ( USNM). Paralectotype: ♂: New York, type USNM 38951 ( USNM).

Tachyusa subaluatacea Casey. Holotype: ♂: North Carolina, Asheville , type USNM 38950 ( USNM).

Additional material. CANADA. British Columbia: 13 mi W Osoyoos , 5.VI.1968, Campbell & Smetana, 3 exx ( CNC) . Saskatchewan: Rt. 3 & North Saskatchewan River , 16 mi N & 16 mi E Lloydminster, 2.VI.1975, L. & N. Herman, 2 exx ( AMNH) . UNITED STATES. Arizona: Yuma , 6.V.1911, A. Fenyes, 2 exx ( BMNH) . Arkansas: Crawford Co., 6–15 mi SSE Winslow (Washington Co.), Bayou Creek , 26.V.1987, L. Herman, 2 exx ( AMNH) ; Newton Co., Buffalo National River, Steel Creek , 17.V.1986, J. M. Campbell, 1 ex ( CNC) . Colorado: Deer Creek , 13.VII.1938, Green, 16 exx ( CAS) ; Huerfano Co., 8 mi SSW La Veta, Cucharas River , 7800 ft., 18.VIII.1982, L. Herman, 3 exx ( AMNH) ; 1 ex ( ISEA). Illinois: Algonquin , 1 ex ( CAS) ; Havana , 11.VIII.1907, J. D. Hood, 1 ex ( CAS) ; Kankakee Co., E of Bourbonnais, Kankakee River , 2.VII.1955, H. Dybas, 2 exx ( FMNH) ; Livingston Co., W of Cornell, Wagner Farm , 25.VII.1967, 10 exx ( MCZ) ; 4 exx ( FMNH); “ West Sprgs. ”, 26.VI.1944, H. Dybas, 2 exx ( FMNH) ; Winnebago, Co., Rockford , 30.VIII.1988, R. Baranowski, 2 exx ( MZLU) . Indiana: Parke Co., 8.5 mi N Rockville, Sugar Creek , 4.VIII.1977, L. & N. Herman, 1 ex ( AMNH) ; Tippecanoe Co., 15.V.1971, N. M. Downie, 3 exx ( FMNH) . Kansas: Lawrence , 8.VII.1933, Blackwelder, 1 ex ( AMNH) ; Onaga , 1 ex ( CAS) ; Tasco , S. Fork Salomon River, Sheridan Co., 3.VIII.1965, 1 ex ( AMNH) . Kentucky: Trigg Co., Port Prizer Point Resort, by Lake Barkley , 12 mi NW Cadiz, 14.VI.1983, B.M. Lysy, 1 ex ( MCZ) . Massachusetts: Blancka , 1 ex ( CAS) ; Lawrence , 2 exx ( FMNH) ; Frederick, Blanchard , 46 exx ( MCZ) ; Northampton , 15.VIII.1970, G.J. Kiteley, 1 ex ( CNC) . Minnesota: Swift Co., 19 mi W Benson, Pomme de Terre River , 31.VIII.1982, L. Herman, 4 exx ( AMNH) ; 1 ex ( ISEA). Montana: Broadwater Co., 2 mi S Townsend , 22.VIII.1970, L. Herman, 2 exx ( AMNH) ; Meagher Co., 23 mi NE White Sulphur Spgs. , 23.VIII.1970, L. Herman, 1 ex ( ISEA) ; Musselshell Co., 38 mi E of Roundup, Musselshell R., 24.VIII.1970, L. Herman, 1 ex ( AMNH) ; Broadwater Co., Townsend, Missouri River , 22.VIII.1970, L. Herman, 4 exx ( AMNH) ; Kalispell, Wickham , 1 ex ( MCZ) ; Rosebud Co., Forsyth, Yellowstone River , 21.VIII.1970, L. Herman, 7 exx ( AMNH) ; 1 ex ( ISEA). Nebraska: Sheridan Co., 14 mi S Hay Springs, Niobrara River , 5.IX.1970, L. Herman, 1 ex ( AMNH) ; Cherry Co., 3 mi NE Valentine, Niobrara River , 23.VIII.1974, R. T. Schuh, 3 exx ( AMNH) ; Garden Co., Oshkosh, North Platte River , 6.VIII.1970, L. Herman, 2 exx ( AMNH) . New York: Ithaca, Inlet Valley , VII.1919, 2 exx ( SEMC) . New Mexico: Doña Ana Co., 1 mi NE Hatch, 4050 ft., shore of Rio Grande , 8.V.1968, L. Herman, 3 exx ( AMNH) ; 1 ex ( ISEA); Sandia Mts., 7000 Cibola National Forest, Las Huertas Creek , 8.VII.1969, Smetana, 1 ex ( CNC) ; 2 exx ( ISEA); Taos Co., Rio Grande State Park , 20.VI.1983, Askevold, 4 exx ( CNC) . New Hampshire: Manchester , 3.VII.1935, J.W. Green, 4 exx ( CAS) . New Jersey: Philipsburg , 25.VII.1915, J.W. Green, 1 ex ( CAS) ; 19.V.1918, 1 ex ( CAS) ; 8.VIII.1933, 11 exx ( CAS) ; 11.VII.1934, 11 exx ( CAS) ; 27.VI.1937, 1 ex ( CAS) ; Sussex Co., Strokes State Forest , 18–19.V.1974, L. & N. Herman, 1 ex ( AMNH) . Ohio: Morgan Co., 8 mi NNW McConnelsville, Muskingum River , 31.VII.1977, L. & N. Herman, 3 exx ( AMNH) ; Muskingum Co., 5 mi SE Zanesville, Muskingum River , 23.IX.1970, L. Herman, 2 exx ( AMNH) . Pennsylvania: Easton , 20.VI.1915, J.W. Green, 24 exx ( CAS) ; Monroe Co., Delaware Water Gap, Delaware River , 29.VI.1971, L. Herman, 4 exx ( AMNH) . South Dakota: Pennington Co., 5–7 mi SSW Hill City, Spring Creek , 5550 ft., 16.VI.1981, L. Herman, 1 ex ( AMNH) ; 1 ex ( ISEA). Texas: Columbus , 2 exx ( USNM) . Vermont: Royalton , 3.VI.1985, E. Kiteley, 2 exx ( CNC) ; Swanton , 28.VI.1991, E. Kiteley, 1 ex ( CNC) . Washington, D.C.: 9.V.1906, 1 ex ( USNM) . Wyoming: Converse Co., Douglas, North Platte River , 4.VIII.1970, L. Herman, 1 ex ( AMNH) ; Fremont Co., 13.5 mi SE Lander Twin Creek , 5500 ft., 13.VI.1981, L. Herman, 6 exx ( AMNH) ; Sheridan Co., 0.7 mi W Leiter, Clear Creek , 3750 ft., 26.VII.1978, L. & N. Herman, 26 exx ( AMNH) ; 1 ex ( ISEA).

Redescription. Body. Length 2.5–2.8 mm, convex, parallel­sided, glossy; head pitchy brown to black, pronotum brown, elytra brownish red to brown, abdomen black with three basal tergites red to reddish brown, legs yellow, antennae brown with antennomeres 1–3 yellowish red.

Head quadrate, flattened dorsally, broadly and relatively shallowly impressed medially in male and without impression in female; eyes large, moderately protruding from lateral contours of head, length of each seen from above subequal to that of postocular region; surface of head with isodiametric mesh microsculpture in male and without microsculpture in female; puncturation fine and moderately dense. Antennae long, clearly increasing in width apically, antennomere 3 shorter than 2, antennomeres 4–7 longer than wide, antennomeres 8–10 quadrate, antennomere 11 nearly conical.

Pronotum transverse, lateral sides gradually narrowed in straight line to obtuse hind angles; before base with small and shallow transverse impression; broadly and relatively shallowly impressed medially in male and without medial impression in female; surface with isodiametric mesh microsculpture in male and without microsculpture in female; puncturation fine, dense and asperate; pubescence at midline directed posteriorly.

Elytra subquadrate, at suture as long as pronotal length at midline; surface lacking microsculpture; puncturation fine, dense and asperate.

Abdomen parallel­sided, bases of tergites 3–5 each with deep transverse impression, impressions with 7–8 longitudinal ridges and distinct median carina, tergal puncturation fine and dense, punctures small and well visible, surface without microsculpture.

Male. Aedeagus as in Figs. 135–136.

Female. Spermatheca as in Fig. 137.

Taxonomic notes. An examination of the types listed above, particularly the genitalia, revealed that they are all conspecific. Therefore, Tachyusa dakotana Casey, 1906 , T. illini Casey, 1906 , T. missouriana Casey, 1906 , T. parviceps Casey, 1906 , T. pruinosa Casey, 1906 and T. subaluatacea Casey, 1906 are here synonymized with Tachyusa obsoleta Casey, 1906 .

Remarks. See under T. cavicollis .

Distribution. Tachyusa obsoleta is widely distributed throughout the United States and also recorded in Canada.