Tachyusa objecta Mulsant & Rey, 1870

Pa ṡnik, Grzegorz, 2006, A revision of the World species of the genus Tachyusa Erichson, 1837 (Coleoptera, Staphylinidae: Aleocharinae), Zootaxa 1146, pp. 1-152 : 106-110

Tachyusa objecta Mulsant & Rey, 1870 ( Figs. 106–108)

Tachyusa objecta Mulsant & Rey, 1870: 183 .

Tachyusa objecta: Mulsant & Rey, 1875: 387 .

Tachyusa (s. str.) objecta: Ganglbauer, 1895: 246 ; Fenyes, 1920: 172; Bernhauer & Scheerpeltz, 1926: 585; Lohse, 1974: 70.

Type material. Lectotype (here designated): ♂: Beaujolais (little circular beige label) ( MHNL) ; Paralectotypes: ♂ and ♀: same data as the lectotype : ♂: Provence, Var (83), Alpes maritimes (06), Belleville ( MHNL) .

Additional material. Austria: Albern near Wien, 1 ex ( NMW) ; Burgenland, Neusiedlersee, Leitha­Aatarm , 30.VIII.1985, 1 ex ( VAPC) ; Donauauen near Klosterneuburg, O. Scheerpeltz, 2 exx ( ETH) ; Donauauen near Albern , Scheerpeltz, 4 exx ( NMW) ; Donauinondation near Wien, W. Stemmer, 1 ex ( NMW) ; Grunburg, Bernhauer , 1 ex ( FMNH) ; Inrersdorf near Wien, Liesingbach river , 28.V.1946, Meyer, 2 exx ( MHNG) ; Kampthal , 3 exx ( NMW) ; Lavant near Wolfsberg , 1 ex ( MIZ) ; Neudorf , 1 ex ( NMW) ; Podersdorf , 16–22.VI.1966, R. Papperitz, 2 exx ( SMNS) ; Sankt Johann near Hohenburg, T. Wanka, 1 ex ( NMW) ; Stockerau, Bernhauer , 2 exx ( FMNH) ; Luze , 1 ex ( NMW) ; Skalitzky , 1 ex ( NMW) ; Ulrichskirchen, Spurny , 1 ex ( IRSNB) ; Waltersdor near Wien, 1 ex ( NMW) ; Wien, Reitter , 1 ex ( HNHM) ; 1 ex ( NMW); Wien, Prater , Breit, 4 exx ( DEI) ; 5 exx ( NMW); Wienerwald , 14.VII.1935, P. Meyer, 1 ex ( BMNH) . Azerbaijan: Lenkoran, Leder, 1 ex ( HNHM) ; 1 ex ( NMW). Belgium: Celles, Lesse river , 19.VII.1938, Fagel, 1 ex ( IRSNB) ; Furfooz, Chaleux , 30.VIII.1946, Fagel, 1 ex ( IRSNB) . Bosnia and Herzegovina: Brcka, V. Zoufal, 4 exx ( NMPC) ; “ Jaice ” [?], 1 ex ( NMW) . Bulgaria: 2 km S Micurin Grabenufer , 1.VII.1979, Uhlig, 1 ex ( ZMHB) ; Burgas, VI.1933, 1 ex ( NMPC) ; Kazanlak, Stausee Georgi Dimitrow , 19.IX.1977, Hieke & Uhlig, 59 exx ( ZMHB) ; Rupite near General Todorow , 4.V.1984, Hieke, 1 ex ( ZMHB) ; Schwarzmeerkuste, Orizare near Nesschar , 12.V.1985, Wrase, 1 ex ( ZMHB) ; Schwarzmeerkuste, Koscharrze , 8.V.1985, Wrase, 1 ex ( ZMHB) ; Verna , VII.1908, Rambousek, 3 exx ( NMPC) . Croatia: Istrien, Pola, F. Lang, 1 ex ( NMW) ; Rjeka , 1 ex ( FMNH) ; 1 ex ( HNHM); 3 exx ( MTD); Zagreb, 1916, Hochetlinger, 1 ex ( NMPC) . Cyprus: Limassol, Limnatis River near Alassa , 300 m, 8.IV.1995, Assing, 4 exx ( VAPC) . Czech Republic: Čelakovice , 15.IV.1917, Rambousek, 1 ex ( NMPC) ; “ Eger ” [Cheb], 1 ex ( IRSNB) ; “ Spindelmuhle ” [Spindleruv Mlyn], Skalitzky, 1 ex ( NMW) . France: Avignon , 17.VIII.1905, Chobaut, 1 ex ( MNHN) ; Adour , 2 exx ( MNHN) ; Bonneuil , VIII.1938, Levasseur, 3 exx ( MNHN) ; Bordeaux , 12.II.1925, G. Tempere, 4 exx ( MHNG) ; 14.IV.1941, G. Tempere, 2 exx ( MHNG) ; Bourges , 20.IV.1911, 2 exx ( MNHN) ; V.1911, 1 ex ( MNHN) ; Chateauroux , 20.VI.1947, Coiffait, 1 ex ( MNHN) ; Corse, Pisciatella , III.1916, 1 ex ( MNHN) ; Corse, Fium d'Orbo, Ghisonaccia , 19–21.IV.1927, 1 ex ( MNHN) ; 3 exx ( NMPC); Corse, Ghisonaccia graviers du Fiumorbo , 2–8.V.1955, G. Fagel, 2 exx ( IRSNB) ; Dax , VI.1906, 2 exx ( MNHN) ; Écully Rhône, G. Audras, 1 ex ( MHNL) ; Gironde, Bordeaux , 19.V.1946, 3 exx ( IRSNB) ; 16.VI.1945, Coiffait, 7 exx ( MNHN) ; Gironde, Cauderan , IV.1951, Coiffait, 1 ex ( MNHN) ; Gironde, Haillan , 1.IV.1946, Coiffait, 1 ex ( MNHN) ; Gironde, Merignac , 7.VIII.1945, Coiffait, 1 ex ( MNHN) ; Grosbois, Côte dOr, VI.1958, 14 exx ( MNHN) ; Indre, Velles , V.1938, Coiffait, 1 ex ( MNHN) ; Côte dOr, V.1949, Levasseur, 5 exx ( MNHN) ; Lyon , 1 ex ( MNHN) ; Morcourt near Kanalufer , 17.VIII.1918, Salchert, 2 exx ( ZMHB) ; Mourillon , 25.IX.1953, 2 exx ( MHNG) ; Perrusson, Mequignon , 1 ex ( CAS) ; Saclay , V.1937, 1 ex ( MNHN) ; V.1942, 1 ex ( MNHN) ; Levasseur , 8 exx ( MNHN) ; Senart , VIII.1938, Levasseur, 1 ex ( MNHN) ; Soissons , 2 exx ( MNHN) ; Saint Dizier , Marne, Deville, 2 exx ( MNHN) ; Saint Florent , Cher, Deville, 1 ex ( MNHN) ; Strasbourg , VII.1952, 1 ex ( MHNG) ; VIII.1952, 1 ex ( MHNG) ; IX.1953, Coiffait, 8 exx ( MNHN) ; Valmy­Masne , 11.VII.1918, 2 exx ( MNHN) ; Vitry , VIII.1941, 3 exx ( MNHN) ; Wissous , V.1942, 5 exx ( MNHN) . Germany: Darmstadt , 19.VI.1954, 1 ex ( MHNG) ; Geesthacht near Hamburg, 21.VIII.1892, Koltze, 4 exx ( DEI) ; Lobau, C. Mandl, 2 exx ( NMW) ; Neudorf , 1 ex ( NMW) ; Siebenburg , 1 ex ( MTD) ; Ulm , 1 ex ( MTD) ; Waltersdorf, Scheerpeltz , 1 ex ( NMW) . Greece: Makedonien, Nomos Serres, Kerkini­See, NE Ende , 1.V.1994, Schawaller, 1 ex ( SMNS) ; Saloniki, Vardareb , 1 ex ( FMNH) ; Valtos, Euboea , IV.1926, Holtz, 1 ex ( FMNH) . Hungary: Ágasegyháza , 18–26.VIII.1952, Bunday, 1 ex ( HNHM) ; Békés, Endröd , 25.V.1980, Ádám, 2 exx ( HNHM) ; Békés, Gyula, Fekete­körös , 90 m, 27.V.1980, Ádám, 1 ex ( HNHM) ; Békés, Sarkad, Fekete­Körös , 90 m, 8.V.1982, Ádám, 9 exx ( HNHM) ; Békés, Szabadkigyós, Nagy­gyop , 90 m, 26.V.1982, Ádám, 1 ex ( HNHM) ; Bükk hegyseg, Nagybérc , 821 m, 8–10.VI.1954, Kaszab & Székessy, 1 ex ( HNHM) ; Bükki National Park, Miskolc, Garadna­völgy , 250 m, 16.IX.1982, Ádám, 1 ex ( HNHM) ; Bükki National Park, Szarvaskö , Eger, 250 m, 17.VII.1984, Hámori, 1 ex ( HNHM) ; Bükki National Park, Varbo, Dobrica­kút , 350 m, 25.VIII.1981, Ádám & Hámori, 2 exx ( HNHM) ; Dömsöd, Apajpuszta , 28.VIII.1952, Kaszab, 5 exx ( HNHM) ; 15.VII.1953, Kaszab, 1 ex ( HNHM) ; Heves, Egerbakta, Baktai­tó , 250 m, 24.IV.1983, Ádám, 3 exx ( HNHM) ; Hortobágy National Park, Hortobágy ­Máta , 10–11.VI.1976, Hámori, 1 ex ( HNHM) ; Hortobágy National Park, Tiszacsege , 22.VIII.1975, Mahunka, 2 exx ( HNHM) ; Hortobágy National Park, Újszentmargita , 16–17.VII.1974, Kaszab, 6 exx ( HNHM) ; Kalocsa, Speiser , 2 exx ( HNHM) ; Kaposfüred, Peregi , 1 ex ( HNHM) ; Kiskunság National Park, Bugac , 12–13.VI.1979, Uhlig, 2 exx ( ZMHB) ; Kiskunsági National Park, Dunapataj, Szelidi­tó , 15.VI.1980, Ádám, 1 ex ( HNHM) ; Kiskunsági National Park, Kiskörös, Szucsi­erdö mocsár, 1.VI.1998, Tóth, 1 ex ( HNHM) ; Kiskunság National Park, Kunfehertó , 6 km SW Kiskunhalas Seeufer, 13.VI.1979, Uhlig, 1 ex ( ZMHB) ; Kiskunsági National Park, Kunfehertó, Feher­tó , 29.VIII.1979, Hámori, 4 exx ( HNHM) ; Kiskunsági National Park, Lakitelek, Tóserdö , Holt­Tisza , 23.V.1977, L. Tóth, 2 exx ( HNHM) ; 26.VII.1977, L. Tóth, 2 exx ( HNHM) ; 20.IX.1978, Hámori, 1 ex ( HNHM) ; Kiskunsági National Park, Pálmonostora, Péteri­tó , 23.VII.1980, Migaly, 1 ex ( HNHM) ; Kiskunság National Park, Tóserdö, 25 km Kecskemét, Holt­Tisza river , 14.VI.1979, Uhlig, 3 exx ( ZMHB) ; Mohács , 1908, Kaufmann, 1 ex ( HNHM) ; Nyírség, Bátorliget , 17–28.VI.1948, Kaszab & Székessy, 1 ex ( HNHM) ; Ócsa, Nagyerdö , 8.VIII.1952, Kovács, 4 exx ( HNHM) ; 24.IV.1952, Kakassné, 1 ex ( HNHM) ; “ Sátoristye ”?, Meschnigg, 2 exx ( MNHN) ; 1 ex ( NMW); Szeged, V. Stiller, 9 exx ( HNHM) ; Velencei­tó, Kis­Velence , 14.IX.1951, Kaszab, 4 exx ( HNHM) . Italy: 50 km W Firenze, Padule di Fucecchio , 16.VI.1992, Assing, 4 exx ( VAPC) ; Basilicata Oasi, WWE "L.S. Giuliano", Cagnolino , 9.VIII.1992, Angelini, 1 ex ( AZPC) ; X.1994, Angelini, 1 ex ( ISEA) ; 1 ex ( VAPC); Episcopia PZ, rive Sinni , 500 m, 22.VII.1993, Zanetti, 1 ex ( AZPC) ; Lombardia, Cava Manara , 16.V.1947, 1 ex ( MSNM) ; Piena del Po, Moncalieri , 16.VI.1912, 1 ex ( MNHN) ; Po river , 16.V.1947, 3 exx ( MCSN) ; Sardegna, Ceresa , V.1935, L. Coghinas, 1 ex ( MSNM) ; Sardegna, Tadesuni, Omodeo , 115 m, 21.V.1995, F. Angelini, 14 exx ( VAPC) ; Sardegna, Oristano , 17.V.1873, R. Gestro, 2 exx ( MCSN) ; Torino, T. Sangone, 1 ex ( MNHN) . Macedonia: Utovo , 1902, Apfelbeck, 2 exx ( MTD) . Norway: Fiume [?], P. Meyer, 3 exx ( NMW) . Poland: Przemyśl , Trella, 1 ex ( ISEA) . Romania: Brad , 2 exx ( MTD) ; Comana Vlasca, A.L. Montandon, 3 exx ( NMPC) ; Galati, V.1902, 9 exx ( BMNH) ; Lacu­Sarat, A.L. Montandon, 1 ex ( NMPC) . Russia: “ Sarepta " [Volgograd], V. Bodemeyer, 3 exx ( FMNH) ; Reitter , 1 ex ( HNHM) ; West Siberia, Barnaul, Ob River , 1918–1920, Frieb, 1 ex ( NMW) . Serbia: Ruma, Hensch, 2 exx ( FMNH) . Slovakia: Komarno , 1946, Havelka, 1 ex ( NMPC) ; Somotor , 21.V.1980, R. Borovec, 3 exx ( VAPC) . Slovenia: Podgrad, Radgona , 28.IV.1998, B. Drovenik, 1 ex ( VAPC) . Spain: Estremadura, Guadalupe , V.1958, Fagel, 8 exx ( IRSNB) ; Madrid, 4 exx ( BMNH) ; Ponferrada, Paganetti , 6 exx ( FMNH) ; Ponferrada, Paganetti , 1 ex ( HNHM) ; Rio Guadalhorce near Alora , 32 km NW Malaga, 3.V.2000, F. Hieke, 2 exx ( ZMHB) ; Rio Escorial , 1 ex ( BMNH) ; Villariciosa, Cordoba , 17.IV.1969, Comellini, 2 exx ( ISEA) ; 21.IX.1971, Comellini, 1 ex ( ISEA) . Tunisia: Ain Draham area , 5–18.V.1988, 1 ex ( ZMUC) . Turkey: Balikesis, Lake Manyas , 1.V.1954, Coiffait, 20 exx ( MNHN) ; Edirne, 8–13.VI.1947, 2 exx ( NMPC) ; Elmali­Antalya, Bey­Daglari , VII.1973, M. & G. Osella, 40 exx ( MSNV) ; Komya, Beysehir , 29.V.1954, Coiffait, 6 exx ( MNHN) ; Tasova­Ladik, Amaays NE, 10.VII.1996, Bayer & Winkelmann, 1 ex ( MSPC) . Ukraine: Dobrowolany near Zaleszczyki, 1 ex ( MIZ) ; Dzwinogród near Borszczów , 27.VII.1937, 1 ex ( FMNH) ; 12.VIII.1937, Tenenbaum, 1 ex ( FMNH) ; Zaleszczyki , 17.IX.1936, Tenenbaum, 1 ex ( FMNH) .

Redescription. Body. Length 2.5–3.3 mm, convex, parallel­sided, glossy; body colour brown to pitchy brown; elytra brownish red, three basal tergites often paler, reddish brow to brown, legs red, antennae reddish brown with antennomeres 1–3 red.

Head circular in outline; convex, eyes large, moderately protruding from lateral contours of head, length of each seen from above subequal to that of postocular region; surface of head without microsculpture; puncturation fine and moderately dense. Antennae long, clearly increasing in width apically, antennomere 3 shorter than 2, antennomeres 4–8 longer than wide, antennomeres 9–10 sexually dimorphic, elongate in male and quadrate in female, antennomere 11 nearly conical.

Pronotum transverse, lateral sides gradually narrowed in straight line to obtuse hind angles; before base with small and shallow transverse impression; surface without microsculpture; puncturation very fine, moderately dense and slightly asperate; pubescence at midline directed posteriorly.

Elytra subquadrate, at suture as long as pronotal length at midline; surface lacking microsculpture; puncturation fine, dense and asperate.

Abdomen parallel­sided, bases of tergites 3–5 each with deep transverse impression, impressions with 7–8 longitudinal ridges, tergal puncturation fine and moderately dense, punctures moderately small and well visible, surface without microsculpture.

Male. Aedeagus as in Figs. 106–107.

Female. Spermatheca as in Fig. 108.

Remarks. Tachyusa objecta is similar to T. nitidula , but differs by the pronotum wider than long, the shorter elytra, the less convex pronotum, the denser pronotal pubescence and particularly by the shape of the genitalia.

Distribution. T. objecta is widespread in the Palaearctic Region, from Spain in the west to the Ob River in the east.