Pseudodryinus burmensis, Olmi & Xu & Guglielmino, 2014

Olmi, Massimo, Xu, Zaifu & Guglielmino, Adalgisa, 2014, Descriptions of new fossil taxa of Dryinidae (Hymenoptera: Chrysidoidea) from Burmese amber (Myanmar), Acta Entomologica Musei Nationalis Pragae 54 (2), pp. 703-714 : 705-706

Pseudodryinus burmensis sp. nov.

( Fig. 1 )

Type material. HOLOTYPE: J( SMNS Bu-106): MYANMAR: specimen in Lower Cenomanian Burmese amber.

Diagnosis. Temple very long (longer than half length of eye ( Fig. 1D )); occipital carina complete; epicnemium exposed; notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of posterior ocellus (1.5: 2); forewing with costal, median and submedian cells clearly enclosed by pigmented veins; tibial spurs 1/1/2. Description. Male ( Fig. 1 ). Fully winged; length 1.8 mm. Forewing longer than mesosoma + metasoma (60: 54). Length of mesosoma + metasoma: 1.3 mm. Colour brown-testaceous, except legs testaceous.Antenna ¿liform, about three times as long as head (55: 19). Antennal hairs very short, much shorter than breadth of antennal segments. Antennal segments in following proportions: 5: 4.5: 6: 6: 6: 6: 5.5: 5: 5: 6. Head dull, apparently reticulate rugose, about twice as long as eye (19: 9). Clypeus very long, ovoid. Mandible very long, with teeth not distinct. Eye normally protruding, covered with very short hairs. Occipital carina complete. Posterior ocelli touching occipital carina. Temple distinct, very long. POL = 3; OL = 2; OOL = 3; TL = 4; greatest breadth of posterior ocelli about as long as OL; frontal line not visible. Propleuron short, forming a neck between head and pronotum, deeply inserted in occiput. Pronotum short, much shorter than head (3: 19) and scutum (3: 9); pronotal tubercle reaching tegula. Scutum apparently bare, apparently unsculptured. Notauli complete, posteriorly separated; minimum distance between notauli shorter than greatest breadth of posterior ocellus (1.5: 2). Scutellum apparently unsculptured, shorter than scutum (6: 9). Metanotum shorter than scutellum (3: 6), apparently unsculptured. Epicnemium exposed. Shape of pronotum, scutum, scutellum and metanotum similar to that of extant males of Pseudodryinus . Propodeum longer than scutum (12: 9), with dorsal surface reticulate rugose, with very large areolae; posterior surface with two complete longitudinal keels and median area apparently unsculptured. Forewing hyaline, without dark transverse bands, with normal venation of extant Pseudodryinus , with three basal cells (costal, median, submedian) clearly enclosed by pigmented veins. Pterostigma narrow, much longer than broad (10: 2); marginal cell open; distal part of stigmal vein much longer than proximal part (11: 6), almost reaching wing’s border; distal part of stigmal vein not S-shaped; shape of forewing similar to that of extant Pseudodryinus . Petiole distinct, much shorter than propodeum (2: 12). Shape, length and breadth of wings similar to those of extant Pseudodryinus . Shape and morphology of body similar to those of extant Pseudodryinus . Foreleg segments in following proportions: 8 (coxa): 6 (trochanter): 13 (femur): 10 (tibia): 8 (tarsal segment 1): 3 (tarsal segment 2): 3 (tarsal segment 3): 3 (tarsal segment 4): 4 (tarsal segment 5). Midleg segments in following proportions: coxa and trochanter not visible: 16 (femur): 12 (tibia): 22 (tarsus). Hindleg segments in following proportions: coxa and trochanter not visible: 15 (femur): 19 (tibia): 22.5 (tarsus). Maxillary and labial palpi not visible. Tibial spurs 1/1/2.

Female. Unknown.

Etymology. Burmensis, - is, - e (adjective); the species is named after Burma, the former name of Myanmar.

Hosts. Unknown.

Remarks. The new species is the second known fossil of Pseudodryinus Olmi, 1991 . The other species is P. parisiensis Peinado, Nel & Waller, 2006 , described from the Earliest Eocene French amber. A comparison of the new species with P. parisiensis is impossible, because the French species has been described on the basis of a female specimen. The sexual dimorphism in Pseudodryinus (and in all Dryinidae ) is so large that female and male are completely different and morphologically not comparable.

Though the mandible teeth are not distinct through the amber (this character is very important to distinguish males of different dryinid subfamilies), the new species has been assigned to the subfamily Dryininae and the genus Pseudodryinus because of the general aspect and the following characters: temple very long (longer than half length of eye ( Fig. 1D )); occipital carina complete; epicnemium exposed; notauli complete; minimum distance between notauli shorter than greatest breadth of posterior ocellus; forewing with three basal cells (costal, median, submedian) clearly enclosed by pigmented veins; tibial spurs 1/1/2. The above combination of characters cannot be found in other Dryinidae .

Figures

Fig. 1. Pseudodryinus burmensis sp. nov. A – habitus in dorsal lateral view. B, C – head and mesosoma in dorsal view. D – habitus in ventral lateral view. Scale bar: A = 1.51 mm; B = 0.65 mm; C = 0.36 mm; D = 1.2 mm.