Montenegrina janinensis (Mousson, 1859)

Feher, Zoltan & Szekeres, Miklos, 2016, Taxonomic revision of the rock-dwelling door snail genus Montenegrina Boettger, 1877 (Mollusca, Gastropoda, Clausiliidae), ZooKeys 599, pp. 1-137 : 45

Taxon classification Animalia Stylommatophora Clausiliidae

Montenegrina janinensis (Mousson, 1859) Fig. 21J, K

Clausilia janinensis Mousson, 1859: 276.

Clausilia (Delima) janinensis - Westerlund 1884: 54-55.

Delima (Albanodelima) janinensis - Wagner 1924: 120.

Montenegrina janinensis janinensis - Zilch 1981: 129. - Nordsieck 2009: 75.

Montenegrina janinensis crassilabris Fauer, 1993: 55-56, plate 1, fig. 7. - Nordsieck 2009: 75.

Diagnosis.

Shell small, light corneous. Lower whorls smooth, upper ones with fine, widely-spaced, indistinct ribs. Neck inflexed, wrinkled-costate. Basal and peripheral crests well visible. Peristome attached, rounded to somewhat angular, with slightly to strongly swollen, reflexed margin. Lamellae superior and spiralis overlap. In front view lamella inferior moderately emerged, broadly-bent subcolumellaris visible. Lunella dorsolateral, mostly separate from the weak basalis. Subclaustralis residual, sulcalis well developed. Anterior plica superior not connected to the lunella complex. Clausilium plate partly visible through the aperture.

Dimensions

(in mm). Hs: 12.2-17.1, Ws: 3.2-4.2 (Perama, HNHM 99560).

Type locality.

"environs de Ianina" = Greece, Epirus, Ioannina [ janinensis ]; Greece, Epirus, Kria near Perama, above the southeastern cemetery, 600 m [crassilabris].

Type material.

"Janina, Epirus", ex Schaefli, ex Mousson, syntypes [ janinensis ] (SMF 176316/6); same locality, ex Boettger, syntypes [ janinensis ] (SMF 176317/3); same locality, ex Shuttleworth, syntypes [ janinensis ] (NMBE 21041/6); Kria near Perama, leg. Fauer, 10.viii.1971, holotype [crassilabris] (SMF 309237), paratypes [crassilabris] (SMF 309238/3, HNHM 39968/3, SMNS-N 9791, ZMH 91624/pl.); same locality, leg. PS, Fauer, 27.ix.1989, paratypes [crassilabris] (NMBE 535019, NMBE 22589/3, NHMW 103731, ZMH 91623/pl.); Kria, above the church, leg. Hausdorf, 31.v.1985, paratypes [crassilabris] (ZMH 91622/pl.); Kria, above the southeastern cemetery, 600 m, leg. Fauer, 22.iv.1988, paratypes [crassilabris] (ZMH 91627/pl.); 1.2 km from Kria toward Kranoula, ex Fauer, 10.viii.1971, paratypes [crassilabris] (ZMH 91625/6); Perama, exit of the stalactite cave, leg. PS, 29.vii.1975 paratypes [crassilabris] (HNHM 67025/4, NMBE 535017/60, ZMH 91626/9).

Other material.

Greece, Epirus, Amfithea–Spothi road N of Lake Pamvotis, 2 km E of Ligkiades junction, 640 m, 39.6802°N, 20.9052°E, leg. ZE, ZF, JG, 24.vi.2013 (HNHM 99559); Perama, near entrance of the cave, 500 m, 39.6947°N, 20.8463°E, leg. ZE, ZF, JG, 24.vi.2013 (HNHM 99560); Mitsikeli Mts, S of Dikórifo, 1070 m, 39.7828°N, 20.8041°E, leg. PS, 23.v.1991 (NMBE); Kria, dry gorge above the cemetery, 660 m, 39.7245°N, 20.8429°E, leg. ZE, ZF, JG, 24.vi.2013 (HNHM 99562); Perama, near the exit of the stalactite cave, 520 m, 39.6971°N, 20.8433°E, leg. ZE, ZF, JG, 24.vi.2013 (HNHM 99561).

Distribution.

Southwestern part and foothills of the Mitsikeli Mts in northwestern Greece. Most of the known occurrences are along the northern shore of the Pamvotis Lake (between Kria and Spothi), but a record from Dikórifo indicates that the species may be wider distributed (Fig. 22).

Remarks.

Although not synonymized formally, Nordsieck (2009) already questioned the distinct taxonomic status of crassilabris, assuming that it might only be a local form of Montenegrina janinensis with thickened peristome. This morphotype is known from Kria and Perama, but its range is not clearly distinct from that of the typical janinensis . Moreover, on the Goritsa Hill at Perama they live in parapatry: at the entrance of the stalactite cave typical janinensis , whereas at the exit (only a few hundred meters farther) the form with the thickened peristome ( Fauer 1993, and also our personal observation).

Based mainly on their small shell size, originally several non-related Montenegrina taxa were described as subspecies of Montenegrina janinensis . These include Montenegrina soosi Erőss & Szekeres, 2006, and some subspecies currently classified with Montenegrina attemsi , Montenegrina dofleini , Montenegrina grammica , Montenegrina hiltrudae , Montenegrina sporadica Nordsieck, 1974, and Montenegrina tomorosi Brandt, 1961.