Coenagrion scitulum ( Rambur, 1842 )

Kosterin, Oleg E., Onishko, Vladimir V., Ilyina, Elena V., Chepurnov, Grigory Yu. & Blinov, Alexander G., 2024, The genus Coenagrion Kirby, 1890 (Odonata: Coenagrionidae) in the Russian part of the Caucasus, Zootaxa 5471 (2), pp. 151-190 : 174

7. Coenagrion scitulum ( Rambur, 1842)

Fig. 11e–k

Reported from: two localities in Krasnodarskiy Kray: the Pslukh River in Sochi Municipality ( Skvortsov 2010) and Lake Krugloe in Novorossiysk Municipality ( Kosterin & Solovyev 2017); Voskhod Estate of Maykop in Adygea Republic ( Shapovalov & Korotkov 2019); unprecisely for Stavropol’skiy Kray ( Skvortsov 2010).

Specimens. Krasnodarskiy Kray: 2 ♂♂ (37681939, 37681905, 42847761, 108801024, 108802758 ( Fig. 11e )), Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 44.853–857 N, 37.391–393 E, 150–189 m a.s.l., 24.06.2019 ; ♂. Novorossiysk Municipality, Lake Krugloe (remnants of a small lake in a lower part of intermontane valley), 44.6768° N, 37.5973° E, 70 m a.s.l. 10.07.2016, O. Kosterin leg. ; ♂, Goryachiy Klyuch Municipality, Fanagoriyskoe village NW environs, ponds, 44.515–516 N, 39.108–109 E, 101–102 m a.s.l., 27.06.2019 ; ♀ (blue morph), Goryachiy Klyuch Municipality , an overgrown pond 1.5 km WNW of Bezymyannoe village, 44.556 N, 39.108 E, 80–83 m a.s.l., 27.06.2019 ; ♂, Apsheronsk District , 0.5 km NE of Tsurevskiy Settlement, a big former quarry, 44.414–416 N, 39.786–789 E, 207–208 m a.s.l., 30.06.2019 . Karachay-Cherkes Republic: ♀ (blue morph), Prikubanskiy District, 3.3 km WSW of Kavkazskiy village, Lake Maloe NW bank, 44.251–254 N, 42.195–200 E, 610 m a.s.l., 5.07.2019 ; 3 ♂♂ (42847508), 3 ♀♀, Khabez District, Inzhichishkho village N margin, big ponds, 44.211–219 N, 41.714–724 E, 578–582 m a.s.l., 6.07.2019 ; 2 ♂♂ (109974039, Fig. 11f ), Malokarachaevsk District, Uchkeken village E environs, a chain of ponds, 43.934 –43.948 N, 42.564– 42.570 E, 887–923 m a.s.l., 9.07.2019 . Republic of Dagestan: ♂, Kazbekovskiy District, 3.5 km ENE of Dylym village, Lake Gorendzhikor , 43.0607 –43.0620 N, 46.6758– 46.6780 E, 730–732 m a.s.l., 20.06.2021 ; ♂ (♂♂, ♀♀, 176987267, 176987387 ( Fig. 11i ), 176987519, 176987626 ( Fig. 11g ), 180162014, 180162157, 182772197, 182772260, 182772305), Buynaksk District, Buynaksk S suburbs, Nizhnee Kazanishche village environs, the Orta Kopyur Reservoir, 42.7835 N, 47.1362 E, 510 m a.s.l., 29.05.2023 ; ♂ (176987151, 176987519, 176987387, 178697738, 180652685, 180652549, 182772530), Buynaksk District, the water reservoir in Khalimbekaul village , 42.852 –42.858 N, 47.130– 47.133 E, 361 m a.s.l., 29.05.2023 ; 2 ♀♀ (166839019, 186507053), Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 43.2003 N, 47.4869 E, 26 m below s.l., 23.05.2023, O. Kosterin et E. Ilyina leg. ; ♀ (blue morph) (99613386, 99613479), Karabudakhkent District , 1.3 km ENE of Kula village, the former second pond (a boggy meadow), 42.6534 –42.6544 N, 47.6117– 47.6130 E, 212–217 m a.s.l., 19.06.2021 ; ♂ (186510393), Sergokala District, 3 km SW of Myurego village, the Inchkheozen’ River upper valley, 42.3946 N, 47.6774 E, 555 m a.s.l., 23.06.2019 .

Additional photographic registrations: Krasnodarskiy Kray: ♂ (108801024), ♀ (blue morph) (42847863, 108804470 ( Fig. 11j )), Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 44.8565 N, 37.3928 E, 150–189 m a.s.l., 25.06.2019 ; ♂ (37681974), Apsheronsk Town N environs, a Pshekha River right oxbow lake, 44.4169 N, 39.7873 E, 30.06.2019 . Republic of Dagestan: 2 ♂♂; 3 ♀♀ (176987741, 176987626 ( Fig. 11g ), 176988062, 176988292 ( Fig. 11k ), 182772530, 182772690, 182772933 ( Fig. 11k ), 189635377), Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 43.1998 –43.2000 N, 47.4862– 47.4867 E, 26 m below s.l., 30.05.2023 ; 2 ♀♀ (186510794, 186510868, 186590193, 186590343 ( Fig. 11h ), Derbent District , 3.5 km NE of Belidzhi village, thickets at a swamp, 41.9048 N, 48.4452 E, 18 m below s.l., 25.05.2023.

Distribution and habitat. Widespread at lower elevations of the Caucasus but never numerous. Breeds in lakes, rarely pools. These damselflies more readily disperse apart from water than their other congeners, occurring at forest margins or even among steppe vegetation.

Figures

FIGURE 11. Coenagrion ornatum (a–d) and C. scitulum (e–k) in different populations in the Russian part of the Caucasus in nature; a–c, e–f—males; d—copula; g–j—females; k—tandems; a—Buynaksk District, Verkhnee Kazanishche village environs, a brook on the Akpeozen’ River valley left slope, 29.05.2023 (180159191); b—Tabasaran District, the Rubas River valley 2.5 km NE of Sirtych village, Dzharag-Khyan Spring, Sirtych, 28.05.2023 (178695635); c—Dagestan, Magaramkent District, Samur Delta Forest, Bil’-Bil’ Kazmalyar village, 25.06.2021 (178693662); d—Rutul District, the Samur River left arm between Kufa and Kala villages, 26.06.2021 (98025103); e, j—Krasnodarskiy Kray, Anapa Municipality, E suburbs of Supsekh settlement, Lake Zmeinoe env., 24.06.2019 (108802758, 108804470, respectively); f—Karachay-Cherkes Rpeublic, Malokarachaevsk District, Uchkeken village E environs, 9.07.2019 (109974039, Fig. 11f); g, k—Dagestan, Makhachkala N suburbs, pools at the Krivaya Balka River left bank in its lowermost reaches, 30.05.2023 (176987626, 182772933, respectively); h—Derbent District, 3.5 km NE of Belidzhi village, thickets at a swamp, 25.05.2023 (186590343); i—Dagestan, Buynaksk District, Nizhnee Kazanishche village environs, a water reservoir, 29.05.2023 (176987387); a, g–i, k—photo by VVO, b–f, j—by OEK. Not to scale.