Philmontis angulatus, Ingrisch, 2022

Ingrisch, Sigfrid, 2022, Revision of the genus Philmontis Willemse, 1966 and description of a new genus Philmontoides gen. nov. from New Guinea (Orthoptera: Tettigoniidae; Conocephalinae), Zootaxa 5182 (2), pp. 101-151 : 122

Philmontis angulatus sp. nov.

Figs. 1B , 3F, 3M , 5H , 6I , 7M–O

Holotype (male): Papua New Guinea: New Guinea (NE), 30 km South of Garaina, elev. 2000 m (8°4’S, 147°9’E), 9.i.1968, leg. J. & M. Sedlacek —depository: Bernice B. Bishop Museum, Honolulu ( BPBM).

Other specimens studied:— 1 female paratype, same data as holotype ( BPBM) .

Diagnosis. The new species differs from P. nigrofasciatus and related species by the shape of the male cerci that are missing concave excavations and have the apical area not bent mediad but continuing posteriorly although little curved and carries at beginning of about apical third two internal projections. That basic shape of the male cerci, P. angulatus sp. nov. shares with P. profusus sp. nov., with which it also shares the outline of the stridulatory file having a long apical area with reduced teeth. It differs from that species by the apical area of the cerci that is shorter and less strongly curved, the dorsal internal projection of the cercus is also shorter and without widened base, and the ventral internal projection is compressed plate-shaped instead of with styliform end. The female differs from that of P. profusus by the subgenital plate which is strongly curved from one side to the other instead of only little upcurved and clearly visible from below, and by the apical spine-like projections standing rather close together instead of being widely separated in the latter species.

Description. Habitus ( Fig. 1B ): medium-sized species with wings little surpassing abdomen and reaching about hind knees or little surpassing middle of ovipositor in females. Face of uniform color, antennal scrobae partly or fully black. Pronotum lateral lobes long and narrow, its deepest point in males just behind mid-length, in females around beginning of apical third; auditory swelling distinct but without humeral sinus. Pronotum elongate, disc with anterior margin faintly concave, posterior margin broadly rounded. Prosternal spines short; mesosternal lobes obtuse; metasternal lobes rounded. Femora with the following number of spines on ventral margins: (1) 4-6 / 6; (2) 5-6 / 2; (3) 9-13 / 0 (n = 2); hind knee lobes with one spine.

Male. Stridulatory file ( Fig. 3M ): total length 0.8 mm; area with countable teeth 0.66 mm with 47 teeth; from base to end of distinct area 0.6 mm with 37 teeth; area with distinctly spaced, including largest teeth 0.55 mm with 33 teeth (n = 1). Tenth abdominal tergite of about same size as preceding tergites, with lateral angles rounded; apical margin wide-roundly excised for about half the length of the tergite. Cerci in nearly basal two thirds cylindrical but behind basal area with a narrow rim along internal surface that suddenly ends before apical area into an angular projection with rounded angle, while the dorsal area continued as a curved, rounded projection with acute spine at tip; before apical constriction from dorsal margin with a straight and stout internal process that carries two acute spinules at end ( Fig. 3F ). Subgenital plate in basal area extended dorsad on both sides, afterward lateral margins converging and shortly behind mid-length, plate divided into a pair of little diverging and setose apical projections that are slightly widening at end and carry a small stylus at apical margin. Titillators flattened, in situ folded; in opened view with basal and apical areas directed laterad; in apical area only moderately widened, before end narrowed and provided at end with a sclerotized disc with very finely serrulate rim; surface of titillators brownish with scattered larger and darker brown spots ( Fig 5H ).

Female. Subgenital plate markedly wider than long, lateral areas strongly curved dorsad; apical margin thickened, carrying in central area a pair of narrow, elongate, spine-like projections curved dorsad; from basal margin, just before both lateral ends of plate with a short and curved projection on dorso-proximal side; along midline of plate with a faint seam ( Figs 7M–O ). Ovipositor moderately long, curved dorsad, with acute tip ( Fig. 6I ).

Measurements (1 male, 1 female).—Body w/wings: male 28, female 30; body w/o wings: male 25, female 24; pronotum: male 8, female 7; tegmen: male 18.5, female 21; hind femur: male 16.5, female 16.5; ovipositor: female 11 mm.

Etymology. The name of the new species refers to the angular ventral projection of the male cercus; from Latin angularis —angular.

Figures

FIGURE 1. Male habitus in dorsal view with left wings spread.—A, Philmontis forcipatus (Willemse, 1966) [Mt Otto]; B, P. angulatus sp. nov. [Garaina]; C, P. spinosus sp. nov. [Bulldog Rd]; D, P. angustus sp. nov. [Mt Wilhelm]; E, P. profusus sp. nov. [Mt Kaindy]; F, P. flexus sp. nov. [Schraderberg]; G, P. nigrofasciatus Willemse, 1966 [Daulo Pass]; H, P. pumilus sp. nov. [Upper Biaru]; I, left tegmen of P. robustus sp. nov. [Bulldog Road]; J, P. murmur sp. nov. [Murmur Pass]; K, P. pandus sp. nov. [Edie Creek].—Scale bars 10 mm.

FIGURE 3. Male abdominal apex with tenth abdominal tergite and cerci in dorsal view (A, C–D, F–G, I–J, L); cerci in lateral view (B); cerci and subgenital plate in lateral view (E); right cercus in dorsal view (H, K); stridulatory file on underside of left tegmen (M).—A–C, Philmontis spinosus sp. nov. [A–B, Upper Watut; C, Mt Missim]; D–E, P. pumilus sp. nov.; F, M, P. angulatus sp. nov.; G, P. profusus sp. nov.; H–I, P. pandus sp. nov. [H, Wau; I, Edie Creek], J–K, P. robustus sp. nov.; L, P. minimus Willemse, 1966.—The arrow in M marks the end of the area with indistinct teeth of the stridulatory file.—Scale bars 1 mm.

FIGURE 5. Male titillators in oblique ventro-apical (A–B, D–E), strict ventro-apical (C, F–I, K–N), and oblique distal view (J).—A, Philmontis nigrofasciatus Willemse, 1966; B, P. angustus sp. nov.; C–D, P. forcipatus (Willemse 1966); E, P. flexus sp. nov.; F, P. spinosus sp. nov.; G, P. profusus sp. nov.; H, P. angulatus sp. nov.; I, P. pumilus sp. nov.; J–K, P. murmur sp. nov.; L, P. minutus Willemse, 1966; M, P. robustus sp. nov.; N, P. pandus sp. nov.—Scale bars 1 mm.

FIGURE 6. Female habitus in lateral view.—A, Philmontis nigrofasciatus Willemse, 1966 paratype; B, P. murmur sp. nov.; C, P. angustus sp. nov. [Mt Wilhelm]; D, P. forcipatus (Willemse, 1966); E, P. extensus sp. nov. [Fly River]; F, P. banz sp. nov. [South of Banz]; G, P. spinosus sp. nov. [Mt Missim]; H, P. profusus sp. nov. [Mt Kaindi]; I, P. angulatus sp. nov. [Garaina]; J, P. flexus sp. nov. [Schraderberg]; K–L, P. pandus sp. nov. [Dowalo]; M, P. pumilus sp. nov. [Upper Biaru|.—Scale bars 10 mm.

FIGURE 7. Female subgenital plate in about ventral view (A, C, F, H, L), oblique ventral view (G, I), lateral view (B, D–E, J–K, M–N), and apical view (O).—A–B, Philmontis nigrofasciatus Willemse, 1966 [Daulo Pass]; C–E, P. angustus sp. nov. [C–D, Mt Wilhelm; E, Keglsugl]; F–H, P. murmur sp. nov. [F–G Murmur Pass; H, Tambul]; I–J, P. banz sp. nov. [Banz]; K–L, P. flexus sp. nov. [Schraderberg]; M–O, P. angulatus sp. nov. [Garaina], M from right, N from left side.—Abbreviations: ap apical projection, ba base of subgenital plate, le lateral extension. The arrows in M and N point at the proximal projections.—Scale bars 1 mm.