Reddyanus furai, Kovařík & Šťáhlavský, 2019

Kovařík, František & Šťáhlavský, František, 2019, Revision of the genus Reddyanus from Southeast Asia, with description of five new species from Cambodia, Malaysia, Thailand and Vietnam (Scorpiones: Buthidae), Euscorpius 295, pp. 1-45 : 4-8

Reddyanus furai sp. n.

( Figures 1–36 , 193 , 204 , 225 , Tables 1 , 4 )

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TYPE LOCALITY AND TYPE REPOSITORY. Vietnam, Pho Yen Province ; FKCP.

TYPE MATERIAL. Vietnam, Pho Yen Province, 2018, 1♂ 1♀ (holotype No. 1453 and paratype), leg. V. Fura, FKCP.

ETYMOLOGY. The species epithet is a patronym honoring VladimÍr Fura, the collector of types of the new species.

DIAGNOSIS. Adult from 33 mm (male) to 37 mm (female) long. Male has longer metasomal segments and telson. Pedipalps, legs and metasoma I–IV yellow, with brown spots. Femur of pedipalps mostly yellow, patella mostly black. Manus of pedipalps yellow with two or three black spots, fingers black. Metasoma V black and telson reddish black. First metasomal segment bears 10 carinae, second through fourth bear eight carinae, lateral carinae on metasoma II represented only by several granules mainly in posterior part. Metasoma V bears five carinae which can be reduced. Terminal tubercle on second and third metasomal segments of male only very slightly enlarged. Telson elongate in male, shorter in female. Subaculear tooth large but not rounded, dorsally with 4–5 granules in three rows. Pectinal teeth number 14 in male, 13 in female.

DESCRIPTION. The adults are 33.68 mm (male holotype) – 37.31 mm (female paratype) long. The habitus is shown in Figs. 1–6 . For position and distribution of trichobothria of pedipalps see Figs. 21–24 and 26–27 . Sexual dimorphism: adult male with longer and narrow metasomal segments and telson.

Coloration ( Figs. 1–6 ). The base color is yellow with dark spots. The chelicera is yellow, strongly reticulated. The fingers of chelicerae are spotted. The carapace and mesosoma are with a pattern that forms three dark longitudinal strips. The ventral side of the mesosoma is yellowish white. The femur and patella of pedipalps are dorsally and laterally yellow with dark spots, femur is mostly yellow and patella mostly black. The legs are also yellow with black spots. The metasoma I–IV is yellowish or redish brown with black spots, metasoma V is black and telson is reddish brown.

Carapace and mesosoma ( Figs. 3–10 ). The entire carapace is covered with large granules; carinae are absent. The anterior margin of the carapace is medially convex. The mesosoma bears one median carina and is strongly granulated. Tergite VII is pentacarinate. The pectinal tooth count is 14 in male holotype and 13 in female paratype. The pectine marginal tips extend to the quorter of the fourth sternite in the male and to the third quorter of the third sternite in the female. The pectines have three marginal lamellae and 10–11 middle lamellae. The lamellae and fulcra bear numerous light setae. Sternites are smooth with two parallel furrows except sternite VII which bears four incomplete carinae and is smooth. The glabrous zone in the middle of the posterior margin of sternite V present in male in shape of triangle.

Metasoma and telson ( Figs. 16–17 , 31–36 ). The first segment bears 10 carinae. The lateral carinae on the second segment are present but incomplete. The third and fourth segments bear eight carinae, and the fifth segment bears five carinae. Ventral carina is present on the telson in male, and ventral and lateral carinae are present on the telson in female. Surfaces between carinae are obviously smooth, with small, sparse granules that become more numerous only on metasoma I. Terminal tubercle on the second and the third metasomal segments of male only very slightly enlarged. The telson elongate in male, shorter in female. Subaculear tooth large but not rounded, dorsally with 4–5 granules in three rows.

Pedipalps ( Figs. 18–30 ). The pedipalps are very sparsely hirsute. The femur bears five granulated carinae and patella bears 7 smooth reduced carinae. The chela is smooth without carinae in the male and visible carinae in the female. The chela and patella are smooth and femur is densely granulated. Movable finger with six and fixed finger with seven rows of granules, both with several accessory granules and 6 external and 5 internal accessory granules.

Legs ( Figs. 11–13 ). The legs without tibial spurs. The tibia and tarsomeres of legs with setae not arranged into bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of <20 spiniform setae on ventral surface. The femur bears 3–4 and patella 5 carinae; both feur and patella are granulated.

Measurements. See Table 1 .

AFFINITIES. The described features distinguish R. furai sp. n. from all other species of the genus. They are recounted in the key. Morphologically closest is R. loebli from Sri Lanka, which differs mainly by the extremely enlarged terminal tubercle on each dorsal carina of the third metasomal segment ( Fig. 34 and fig. 366 in KovařÍk et al., 2016: 75). In the region only two species with elongate metasomal segments in males has dark pedipalp patella ( Figs. 193 and 196 ). These two species we can easily distinguish by shape of telson in male which is not so enlarged in R. krasenskyi from Java as in R. furai sp. n. from Vietnam. Telson length/depth ratio in male is 3.43 in R. furai sp. n. and 2.96 in R. krasenskyi .

DISTRIBUTION. Vietnam ( Fig. 225 ).

Figures

Figure 225. Map showing confirmed distribution of Reddyanus spp. in Southeast Asia.

Figures 1–2. Reddyanus furai sp. n., male holotype (1) and female paratype (2) in vivo habitus.

Figures 3–6: Reddyanus furai sp. n. Figures 3–4. Male holotype in dorsal (3) and ventral (4) views. Figures 5–6. Female paratype in dorsal (5) and ventral (6) views. Scale bars: 10 mm.

Figures 7–17: Reddyanus furai sp. n. Figures 7, 9, 11–13, 16. Male holotype, carapace and tergites I–III (7), coxosternal area and sternites III–V (9), left legs II–IV, retrolateral aspect (11–13), and telson lateral (16). Figures 8, 10, 14–15, 17. Female paratype, carapace and tergites I–III (8), coxosternal area and sternites III–V (10), right chelicera dorsal (14) and ventral (15) views, and telson lateral (17).

Figures 18–36: Reddyanus furai sp. n. Figures 18–19, 29–33. Female paratype, pedipalp chela dorsal (18) and external (19) views, movable finger (29) and fixed finger (30) dentition, metasoma and telson in lateral (31), dorsal (32) and ventral (33) views. Figures 20–28, 34–36. Male holotype, pedipalp chela dorsal (20), external (21) and ventral (22) views, patella dorsal (23), external (24) and ventral (25) views, femur and trochanter internal (26), dorsal (27), and ventral (28) views, metasoma and telson in lateral (34), dorsal (35) and ventral (36) views. Trichobothrial pattern is indicated in Figures 21–24 and 26–27. Scale bar: 10 mm (31–36).

Figures 192–202: Reddyanus, males, pedipalp trochanter and femur, patella, and chela dorsal, metasoma II–III in lateral views. Figure 192. R. feti, holotype. Figure 193. R. furai sp. n., holotype. Figure 194. R. hofereki sp. n., holotype. Figure 195. R. jendeki, holotype. Figure 196. R. krasenskyi, Java, Ujung Kulon. Figure 197. R. majkusi sp. n., holotype. Figure 198. R. neradi, holotype. Figure 199. R. petrzelkai, holotype. Figure 200. R. rolciki sp. n., holotype. Figure 201. R. schwotti sp. n., Thailand, Khao sok. Figure 202. R. zideki, paratype.

Figures 203–214: Reddyanus, males, telson lateral. Figure 203. R. feti, holotype. Figure 204. R. furai sp. n., holotype. Figure 205. R. hofereki sp. n., holotype. Figure 206. R. jendeki, holotype. Figure 207. R. krasenskyi, Java, Ujung Kulon. Figure 208. R. majkusi sp. n., holotype. Figure 209. R. neradi, holotype. Figure 210. R. petrzelkai, holotype. Figure 211. R. rolciki sp. n., holotype. Figure 212. R. schwotti sp. n., holotype. Figure 213. R. schwotti sp. n., paratype from Thailand, Khao sok. Figure 214. R. zideki, paratype.