Eotrichocera (Archaeotrichocera) longensis, Dong, Fei, Shih, Chungkun & Ren, Dong, 2014

Dong, Fei, Shih, Chungkun & Ren, Dong, 2014, Two new species of Trichoceridae from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China, ZooKeys 411, pp. 145-160 : 147-150

Eotrichocera (Archaeotrichocera) longensis sp. n.

Etymology.

“longensis” is a Latin word, referring to the long leg of this specimen.

Diagnosis.

Sc rather short about 0.65 times as long as the wing and ending at anterior margin proximad of R2; Rs forking proximad of 2/3 (at about 0.55) times of wing length; the d-cell narrow and long (about 2.5 times as long as wide); A2 short and bending sharply toward anterior margin (angle about 128°).

Holotype.

An almost complete female specimen with well-preserved body, wings and head. Specimen number CNU-DIP-NN2013133. Wing length 9.0 mm, width 3.8 mm (Figs 1A, 2, 3A).

Paratype.

A female specimen with body and wings, specimen number CNU-DIP-NN2013131. Wing length 7.7 mm, width 3 mm (Figs 1B, 3B).

Locality and horizon.

Jiulongshan Formation, Late Middle Jurassic, Daohugou Village, Ningcheng County, Inner Mongolia Autonomous Region, China.

Description.

Based on Holotype, different characters of the paratype CNU-DIP-NN2013131 in brackets. Medium-sized winter crane flies, body length (including head) 13 mm with well preserved wings, body and head, [paratype body length (excluding head) 10.5 mm].

Head: Antenna very long, about 3.5 times as long as the head length, palpi about two times as long as the head length, compound eyes preserved.

Thorax: Much higher, in lateral view, than that of the abdomen, subcircular in shape, with robust and well-developed mesonotum. The halters spoon-type and the length of halters as long as thorax.

Wings: Wing is shorter than abdomen, not covering the end of the abdomen. Length 9.0 mm [Paratype with wing length of 7.7 mm], narrow and long (about 2.5 times as long as wide); venation clear, Sc rather short about 0.68 times as long as the wing [Paratype Sc rather short, about 0.65 times as long as the wing] and ending at anterior margin proximad of R2; crossvein sc-r locating at 2/3 of Rs; Rs arising about one-fifth from the base of the wing; R2+3 about 0.8 times as long as R2+3+4; R2 about one-tenth of length of R3; R3 almost three times as long as the R2+3; dM1+2 0.6 times as long as mM1+2, while M1 2.5 times of the dM1+2; a well developed m-m crossvein about three-fourth length of bM3, closing the d-cell and nearly 0.3 length of d-cell; bM1+2 nearly 1.0 times as long as the length of the r-m and the latter at one-fifth of the d-cell; d-cell narrow and long (about 2.5 times as long as wide) and almost one-fifth length of wing; both crossveins m-m and m-cu intersecting with M4 at the same point; Cu long, curved (angle about 135°) and reaching the wing posterior margin at 0.6 from the base of the wing; the stem of A divided into A1 and A2; A1 long, slightly curving and reaching the wing posterior margin; A2 short, 0.15 (right wing) [Paratype 0.14] times as long as wing and almost 0.3 times as long as length of A1, bending sharply (angle about 128°) and reaching the wing posterior margin.

Legs: Legs slender and long; the hind leg nearly 1.2 times as long as the abdomen and 1.3 times as long as the wing. Tarsus with five segments; the first segment of tarsus (t1) is 1.2 times as long as t2 in mid leg.

Abdomen: Abdomen relatively long and thin, with ten segments. Female genital discernible.

Remarks.

Eotrichocera (Archaeotrichocera) longensis sp. n. is assigned to Thrichocerinae based on the following characters: d-cell medium, m-cu present; A2 short, antennae long, flagellomeres thin, much longer than two times of the head length. It belongs to Eotrichocera (Archaeotrichocera) because of wing length from 7.7 to 9.0 mm and d-cell almost one-fifth of wing length. In addition, it differs from all other known Thrichocerinae by its A2 rather short and bending sharply toward anterior margin (angle about 128°), R2 relatively long, Sc forking proximad of 2/3 (at about 0.55) times of wing length, and d-cell narrow and long. To compare the key characters among the new species and other species of Eotrichocera (Archaeotrichocera) , we set up the Table 2.