Raylilia, Galil, 2001

Galil, Bella S., 2001, A new genus and species of leucosiid crab (Crustacea, Decapoda, Brachyura) from the Indo-Pacific Ocean, Zoosystema 23 (1), pp. 65-75 : 66-67

Genus Raylilia n. gen.

TYPE SPECIES. — Arcania gracilipes Bell, 1855 , by present designation.

OTHER SPECIES. — Arcania mirabilis ( Zarenkov, 1969) ; Arcania uenoi Takeda, 1995 ; Raylilia coniculifera n. gen. n. sp.

ETYMOLOGY. — The genus is named after Dr Ray Manning and his wife, Lili Manning.

DIAGNOSIS. — Carapace globose, rounded. Dorsal surface of carapace bearing obtuse or conical tubercles, lateral margin of carapace denticulate, posterior margin tridenticulate. Intestinal region inflated, demarcated by shallow grooves. Front prominent, bilobed. Orbital margin trisutured. Postorbital region concave. Basal antennular segment sealing antennular fossa. Basal antennal segment inserted in inner orbital hiatus. Anterior margin of efferent branchial channel projecting, medially fissured, visible from in dorsal view. Third maxilliped exognath slightly shorter, more slen- der than endognath, merus half as long as ischium.

DESCRIPTION

Chelipeds long, slender, fingers long, more than half as long as chela, cutting edges unevenly spinulate, tips cross. Pereiopods slender, short, dactyls granulate, setose.

Abdominal sulcus deep, nearly reaching buccal cavity. Male abdomen with third to sixth segments fused, narrowing distally, bearing preapical tubercle, telson one fourth as long as fused segments. Adult female abdomen with fourth to sixth segments fused, greatly swollen.

Male first pleopod with stout basal stalk, flanged on interior margin, bearing slender, setose appendix distally on external margin; tip lamellate, flared. Male second pleopod short, distally scoop-like.

REMARKS

Arcania Leach, 1817 has rounded, pyriform or rhomboidal carapace; its dorsal surface is granulate, spinulate, or tuberculate, and the intestinal region bears a single spine or tubercle. The posterior margin of the carapace is bispinose or bidenticulate. The front is bilobed and uptilted. The antennules are obliquely folded, and do not close off the antennular fossa. The antennular basal segment is lodged in the orbital hiatus. The outer orbital margin is trifissured, the inner orbital margin cleft; the infraorbital lobe is spiniform, prominent, and fused with the bidentate anterior margin of the efferent branchial channel. Third maxilliped endopod bears a vertical row of setae in females. The chelipeds are slender and elongate. The male abdomen has segments third to fifth fused, lacking a preapical tubercle; the adult female abdomen is greatly swollen, segments fourth to sixth fused, the telson laciniate. The first male pleopod is slender and elongate, sinuous or straight.

Raylilia n. gen. is distinguished from Arcania Leach, 1817 in having basal antennular segment entirely sealing the antennular fossa, the anterior margin of the efferent branchial channel medially fissured and separated from orbital margin by a deep groove, the posterior margin of the carapace tridenticulate, third to sixth segments of the male abdomen fused and bearing a preapical tubercle, and the first male pleopod distally expanded, lamellate.