Scaphioides minuta (Chamberlin and Ivie)

Platnick, Norman I., Berniker, Lily & Kranz-Baltensperger, Yvonne, 2012, The Caribbean Goblin Spider Genera Scaphioides and Hortoonops (Araneae, Oonopidae), American Museum Novitates 2012 (3751), pp. 1-64 : 14-31

Scaphioides minuta (Chamberlin and Ivie)

Figures 1–74

Stenoonops minutus Chamberlin and Ivie, 1935: 8 , figs. 1, 2, 14 (male holotype from Tampa, Hillsborough Co., Florida, in AMNH; examined).– Chickering, 1969b: 160, figs. 40, 41 (first description of female).

Scaphioides minuta: Platnick and Dupérré, 2010: 9 .

DIAGNOSIS: The combined presence of a laterally (but not medially) pitted sternum (figs. 62, 66) and a carapace lacking tubercles but bearing pits with reticulating ridges (fig. 61) is diagnostic.

MALE (PBI_OON 38032, figs. 1–29, 61–64, 69–73): Total length 1.22. Elevated portion of pars cephalica smooth, sides strongly reticulate. ALE separated by their radius to diameter. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with continuous transverse groove. Endites with rounded tip bearing short, strong, posterolaterally directed seta. Embolus with proximal portion shorter than distal portion, smooth prolaterally, ridged retrolaterally.

FEMALE (PBI_OON 38932, figs. 30–60, 65–68, 74): Total length 1.34. Anterior genitalic projection relatively short, wide, on smoothly convex base.

MATERIAL EXAMINED: UNITED STATES: Florida: Alachua Co. : Gainesville , Feb. 1–14, 1979, pitfall, oak-pine (E. Becker, CNC PBI_OON 38175), 1♀ ; Newnan’s Lake , 29°39′N, 82°15′W, June 13, 1935 (W. Gertsch, AMNH PBI_OON 21037), 2♂, 1♀ , same (W. Ivie , AMNH PBI_OON 21034), 3♂, 1♀ , Mar. 28, 1957 (W. Gertsch, R. Forster, AMNH PBI_OON 21029, 37318), 2♂, 7♀ , Dec. 21, 1962 (W. Ivie, AMNH PBI_OON 38031), 1♀ ; W Newnan’s Lake , Feb. 12, 1942 (W. Ivie, AMNH PBI_OON 21044), 1♂, 1♀ ; San Felasco State Park, Gainesville , May 1, 2008, leaf litter (X. Wang, AMNH PBI_OON 31066), 2♂, 2♀ . Collier Co.: N Naples , 26°14′N, 81°47′W, Dec. 18, 1962 (W. Ivie, AMNH PBI_OON 1058), 2♀ . Columbia Co. : 12 mi N Lake City , 30°20′N, 82°36′W, Dec. 22, 1962 (W. Ivie, AMNH PBI_OON 38030), 3♂, 1♀ . Hernando Co.: no specific locality, Mar. 23, 1947 (H. Wallace, AMNH PBI_OON 38033), 1♂, 1♀ . Highlands Co.: Archbold Biological Station , 27°11′N, 81°21′W, Apr. 1956, Quercus chapmani litter (C. Hoff, AMNH PBI_OON 38028), 1♀ , Dec. 19, 1962 (W. Ivie, AMNH PBI_OON 37333), 1♂, 1♀ , July 3, 1978, litter, elev. 20 m (J., F. Murphy, AMNH PBI_OON 36820), 1♂ ; Highlands Hammock State Park , Feb. 16, 1970, pitfall, sand pine (M. Muma, H. Greene, FSCA PBI_OON 21214), 1♀ ; Parker Islands , Apr. 1956, bromeliads on fallen maple tree (C. Hoff, AMNH PBI_OON 38029), 1♂ ; Sebring Airport , Feb. 19–20, 1982, fossil dune pine litter (G. Edwards, FSCA PBI_OON 21215), 2♀ . Hillsborough Co.: Tampa, Aug. 26, 1933, among dead leaves on ground (W. Ivie, AMNH PBI_OON 336), 1♂ (holotype). Lee Co. : Fort Myers , winter 1934 (W. Barrows, AMNH PBI_OON 37069), 1♂ . Leon Co.: S Chaires on U.S. 27, July 17, 1965, sawdust pile at edge of cypress swamp (W. Suter, FMNH 71726 , PBI_ OON 43494), 1♀ ; 1 mi W Tallahassee on Fla. Rt. 20 just south of intersection with US 90, July 1, 1965, Berlese, litter (W. Suter, FMNH PBI_OON 10502), 1♂ ; Tall Timbers Research Station , June 21, 1968, pitfall, abandoned field (W. Baker, AMNH PBI_OON 377), 1♀ , Aug. 5, 1968, pitfall (W. Baker, AMNH PBI_OON 375), 3♂ , same, pitfall under cedar in middle of lawn (W. Baker, AMNH PBI_OON 376), 2♂ . Liberty Co.: Torreya State Park , 30°33′N, 84°57′W, Apr. 4, 1957 (W. Gertsch, R. Forster, AMNH PBI_OON 21030), 2♂ , Dec. 18, 1967 (W. Ivie, AMNH PBI_OON 21124), 2♂, 1♀ . Marion Co.: Ocala National Forest , Nov. 28, 1972 (R. Kaplan, FMNH PBI_OON 38340), 3♂ . Martin Co.: Jonathan Dickson State Park , 26°59′N, 80°08′W, Dec. 12, 1962 (W. Ivie, AMNH PBI_ OON 1100), 1♀ ; 8 mi NNW Stuart , 27°17′N, 80°17′W, Dec. 12, 1962 (W. Ivie, AMNH PBI_OON 38032), 2♂, 3♀ . Monroe Co.: Big Pine Key , E end, 24°40′N, 81°22′W, Dec. 13, 1962 (W. Ivie, AMNH PBI_OON 37335), 1♂ ; Big Pine Key, nature trail off Key Deer Blvd. , 24°42.558′N, 81°23.181′W, hand sifting (P. Sierwald, FMNH PBI_OON 38339), 1♂ ; Big Torch Key , June 5–Aug. 28, 1986, hammock forest malaise flight intercept trap (S., J. Peck, AMNH PBI_OON 43549), 1♂ ; Key Largo Key , 25°10′N, 80°20′W, Dec. 17, 1962 (W. Ivie, AMNH PBI_OON 37331), 2♂, 1♀ ; Key Vaca, Marathon , Aug. 7, 1972, Berlese, palm-hardwood litter (S. Peck, AMNH PBI_OON 31176, 37521), 1♂, 1♀ ; Lignum Vitae Key , July 1981, Malaise trap trough (S. Peck, CNC 38176), 1♂, 2♀ ); No Name Key , Aug. 28–Dec. 13, 1986, forest malaise flight intercept trough (S., J. Peck, AMNH PBI_OON 37325), 1♂ . Orange Co.: 3 mi NW Maitland , 28°38′N, 81°24′W, Dec. 10, 1962 (W. Ivie, AMNH PBI_OON 37327), 6♂, 9♀ . Pinellas Co.: Dunedin , 1927 (W. Blatchley, MCZ 68303, PBI_ OON 27173), 1♀ . Polk Co.: Winter Haven , Mar. 29, 1968, pitfall, sand-pine dune (M. Muma, H. Greene, FSCA PBI_OON 21213), 1♀ , Mar. 13, 1969, pitfall, sand-pine dune (K. Stone, FSCA PBI_OON 21211), 1♂ , Sept. 30, 1970, pitfall, sand-pine (M. Muma, H. Greene, FSCA PBI_OON 21212), 1♀ . Putnam Co.: Welaka , May 9, 1949 (A. Van Felt, MCZ 68302, PBI_OON 27172), 1♂ (missing palps) . Sarasota Co. : Myakka River State Park , 27°14′N, 82°16′W, Dec. 26, 1963 (J., W. Ivie, AMNH PBI_OON 21035), 1♀ . Volusia Co. : 2 mi S Orange City , 28°53′N, 81°17′W, Dec. 9, 1962 (W. Ivie, AMNH PBI_OON 37334), 2♂, 2♀ . Georgia: Chatham Co.: 3 mi SE Savannah , 32°04′N, 81°04′W, Apr. 4–May 3, 1943 (W. Ivie, AMNH PBI_OON 1141, 1143, 37328, 37336) ,

FIGS. 79–91. Scaphioides campeche , new species, male (79, 80, 82–86, 88, 89) and female (81, 87, 90, 91). 79, 84. Carapace, dorsal view. 80, 81. Sternum, ventral view. 82, 88. Left embolus, prolateral view. 83, 89. Same, retrolateral view. 85. Left palp, prolateral view. 86. Same, retrolateral view. 87. Epigastric region, ventral view. 90. Genitalia, ventral view. 91. Same, dorsal view.

10♂, 9♀; Savannah Beach , 31°60′N, 80°50′W, Dec. 5, 1962 (W. Ivie, AMNH PBI_OON 1144) , 1♀; 3 mi NW Savannah Beach , 32°01′N, 80°53′W, Mar. 14, 1954 (W. Ivie, AMNH PBI_OON 1146) , 10♂, 11♀. Emanuel Co.: N Swainsboro , 28°53′N, 81°17′W, Dec. 23, 1962 (W. Ivie, AMNH PBI_OON 1142) , 2♀. Glynn Co.: Jekyll Island Park, 31°06′N, 81°25′W, Dec. 6, 1962 (W. Ivie, AMNH PBI_OON 37330) , 1♂. Randolph Co.: outside Grier’s Cave , 10 mi N Cuthbert, Oct. 22, 1941 (A. Archer, AMNH PBI_OON 21031, 21039) , 2♀. Thomas Co.: 8 mi SW Meigs , 30°59′N, 84°11′W, Dec. 18, 1967 (W. Ivie, AMNH PBI_OON 37332) , 1♂. North Carolina: Carteret Co.: Bogue Banks , Oct. 15, 1951 (R. Barnes, AMNH PBI_OON 21041) , 1♂, June 6, 1952 (R. Barnes, MCZ 68301, PBI_OON 27174) , 2♂.

DISTRIBUTION: Southeastern United States (North Carolina, Georgia, and Florida) .

Scaphioides campeche , new species

Figures 79–91

TYPES:Male holotype, female allotype, plus one male and one female paratypes from Campeche, Campeche, Mexico (July 5–6, 1951; L. Stannard), deposited in AMNH (PBI_OON 1380) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. gertschi , S. siboney , and S. hoffi in having sternal pits only around the furrows (figs. 80, 81) and both tubercles and pits with reticulating ridges on the carapace (fig. 79), but differ in having the carapace tubercles restricted to the posterior declivity (fig. 79).

MALE (PBI_OON 1380, figs. 79, 80, 82–86, 88, 89): Total length 1.07. Elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with continuous transverse groove. Endites with rounded tip bearing two short, strong setae. Embolus resembles that of S. minuta but proximal portion shorter, thumblike.

FEMALE (PBI_OON 1380, figs. 81, 87, 90, 91): Total length 1.16. Anterior genitalic projection relatively long, narrow, curved, on triangular base.

OTHER MATERIAL EXAMINED: MEXICO: Yucatán: Chichén Itzá (C. Goodnight, AMNH PBI_OON 37329), 1♂ .

DISTRIBUTION: Mexico (Campeche, Yucatán) .

Scaphioides irazu , new species

Figures 92–100

TYPE: Female holotype from Volcán Irazu, Cartago, Costa Rica (July 27, 1965; A. Chickering), deposited in MCZ (72091, PBI_OON 26486) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. phonetus , S. econotus , and S. nitens in having many tubercles on the carapace, arrayed in triangular patches opposite the coxae (fig. 92), but have a white, rather than patterned or iridescent abdomen and in having the anterior genitalic process situated on a relatively wide base (figs. 99, 100).

MALE: Unknown.

FEMALE (PBI_OON 26486, figs. 92–100): Total length 1.37. Elevated portion of pars cephalica smooth, sides granulate. ALE separated by their radius to diameter. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with interrupted transverse groove. Anterior genitalic process very narrow, on boat-shaped base.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Costa Rica (Cartago).

Scaphioides bimini , new species

Figures 75–78

TYPE: Male holotype from South Bimini, Bahama Islands (May 1951; W. Gertsch, M. Cazier), deposited in AMNH (PBI_OON 21048) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. camaguey , S. cobre , and S. yateras in having large pits widely distributed on the sternum, including along the midline (fig. 76), but differ in having a relatively short, wide embolus (figs. 77. 78).

MALE (PBI_OON 21048, figs. 75–78): Total length 1.17. Elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture covering entire surface, furrows with rows of small pits, anterior margin with interrupted transverse groove. Embolus without clearly distinguishable proximal and distal lobes.

FEMALE: Unknown.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Bahama Islands (South Bimini) .

Scaphioides gertschi , new species

Figures 101–113

TYPES: Male holotype, female allotype, and two male paratypes from South Bimini, Bahama Islands (May 1951; W. Gertsch, M. Cazier), deposited in AMNH (PBI_OON 21008) .

ETYMOLOGY: The specific name is a patronym in honor of Willis Gertsch, one of the collectors of the type series.

DIAGNOSIS: Members of this species resemble those of S. campeche , S. siboney , and S. hoffi in having sternal pits only around the furrows (figs. 102, 103) and both tubercles and pits with reticulating ridges on the carapace (fig. 101), but have the proximal lobe of the embolus rounded in prolateral view (fig. 104) and a long, narrow tip on the anterior genitalic process (figs. 112, 113).

MALE (PBI_OON 21042, figs. 101, 102, 104–108, 110, 111): Total length 1.18. Elevated portion of pars cephalica smooth except posterior margin with U-shaped band of tubercles; carapace sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with continuous transverse groove. Endites with rounded tip. Embolus with proximal portion short, narrow, distal portion arched.

FEMALE (PBI_OON 21042, figs. 103, 109, 112, 113): Total length 1.37. Anterior genitalic projection long, wider distally than proximally, on short base with squared sides.

OTHER MATERIAL EXAMINED: WEST INDIES: Bahama Islands: North Bimini, Dec. 4, 1952 (A. Nadler, AMNH PBI_OON 21032), 2♀; Pigeon Cay, Andros Island , May 1–6, 1994, Berlese, coastal coppice litter (R. Anderson, AMNH PBI_OON 1766), 1♀ ; South Bimini , May 1951 (W. Gertsch, M. Cazier, AMNH PBI_OON 21042, 21045, 43633), 10♂, 1♀ , June 1951 (M. Cazier, C., P. Vaurie, AMNH PBI_OON 21036, 21040), 4♂, 1♀ , July 1951 (C., P. Vaurie, AMNH PBI_ OON 21046), 1♀ , Apr. 10–14, 1952 (E. Mayr , AMNH PBI_OON 21047), 1♂ , Dec. 12–18, 1952 (A. Nadler, AMNH PBI_OON 21038, 21043), 3♂, 1♀ , Mar. 22–28, 1953 (A. Nadler , AMNH PBI_OON 21033), 1♂ , May 26–29, 1953 (A. Nadler, AMNH PBI_OON 21049), 1♀ .

DISTRIBUTION: Bahama Islands (Andros, North Bimini, South Bimini) .

Scaphioides camaguey , new species

Figures 114–126

TYPES: Male holotype, female allotype, plus one male and four female paratypes taken during the dry season in a semideciduous forest at Sierra de Cubitas , Limones de Tuabaquey, Camagüey, Cuba (Feb. 2010; Y. Martínez), deposited in MNH (PBI_OON 968) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. bimini , S. cobre , and S. yateras in having large pits widely distributed on the sternum, including along the midline (figs. 115, 116), but have a relatively long, narrow embolus with a greatly reduced proximal lobe (figs. 117, 118) and a relatively long, narrow anterior genitalic process (figs. 125, 126).

MALE (PBI_OON 968, figs. 114, 115, 117–122): Total length 1.03. Elevated portion of pars cephalica strongly reticulate, sides strongly reticulate, posterior margin of elevated portion with U-shaped band of tubercles. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with interrupted transverse groove. Proximal portion of embolus tiny, distal portion strongly arched.

FEMALE (PBI_OON 968, figs. 116, 123–126): Total length 1.28. Anterior genitalic process much wider posteriorly than anteriorly, on short base with angular edges that extend farther anteriorly than do paramedian portions of base.

OTHER MATERIAL EXAMINED: WEST INDIES: Cuba: Camagüey: Sierra de Cubitas, Limones de Tuabaquey, Sept. 2009 (Y. Martínez, AMNH PBI_OON 969), 2♂, 3♀ .

DISTRIBUTION: Cuba (Camagüey).

Scaphioides granpiedra , new species Figures 127–136 TYPES: Male holotype, female allotype, and female paratype taken in litter at an elevation of 1200 m at a site 7.5 km on the road to La Gran Piedra, 20°00′23″N, 75°40′31″W, Santiago

FIGS. 147–159. Scaphioides cobre , new species, male (147, 148, 150–155) and female (149, 156–159). 147, 152, 156. Carapace, dorsal view. 148, 149. Sternum, ventral view. 150. Left embolus, prolateral view. 151. Same, retrolateral view. 153. Left palp, prolateral view. 154. Same, retrolateral view. 155. Mouthparts, ventral view. 157. Epigastric region, ventral view. 158. Genitalia, ventral view. 159. Same, dorsal view.

FIGS. 168–177. Scaphioides cletus (Chickering) , male (168, 169, 171–174) and female (170, 175–177). 168. Carapace, dorsal view. 169, 170. Sternum, ventral view. 171. Left embolus, prolateral view. 172. Same, retrolateral view. 173. Left palp, prolateral view. 174. Same, retrolateral view. 175. Epigastric region, ventral view. 176. Genitalia, ventral view. 177. Same, dorsal view.

FIGS. 178–190. Scaphioides hoffi (Chickering) , male (178, 179, 181–187) and female (180, 188–190). 178, 183. Carapace, dorsal view. 179, 180. Sternum, ventral view. 181, 186. Left embolus, prolateral view. 182, 187. Same, retrolateral view. 184. Left palp, prolateral view. 185. Same, retrolateral view. 188. Epigastric region, ventral view. 189. Genitalia, ventral view. 190. Same, dorsal view.

de Cuba, Cuba (May 8, 2010; N. Platnick, A. Sánchez, A. Pérez, G. Alayón), deposited in BSC (PBI_OON 960) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. reducta in having relatively few, small sternal depressions that are “haloed” by distinct edges (figs. 128, 129) but have the proximal portion of the embolus no longer than the distal portion (figs. 130, 131) and a smaller base of the anterior genitalic process (figs. 135, 136).

MALE (PBI_OON 961, figs. 127, 128, 130–133): Total length 1.33. Elevated portion of pars cephalica strongly reticulate, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with small round pits, microsculpture everywhere but front, furrows with rows of small pits, anterior margin with continuous transverse groove. Endites with rounded tip. Proximal portion of embolus separated from distal portion, on retrolateral side, by short, round notch.

FEMALE (PBI_OON 361, figs. 129, 134–136): Total length 1.50. Anterior genitalic process hourglass shaped.

OTHER MATERIAL EXAMINED: WEST INDIES: Cuba: Santiago de Cuba: Municipio Santiago de Cuba: Cañón del Río Juticí, Reserva Ecológica Siboney-Juticí , 19°57′44″N, 75°44′48″W, May 4–5, 2010, Berlese, litter, elev. 10 m (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_ OON 963), 8♂, 2♀ (including Eva Guablomme DNA sequencing voucher) ; El Cobre, 20°02′48″N, 75°56′20″W, May 7, 2010, Berlese, litter (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 961), 1♂, 1♀ ; Mata Yegua, 2 km NE El Escandel, near tributary of Río Escandel , 20°05′34″N, 75°41′49″W, May 10, 2010, litter (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 962), 1♂ ; Reserva Ecológica Siboney-Juticí , 19°57′39″N, 75°42′52″W, May 5, 2010, dry litter (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 43634), 3♂, 3♀ .

DISTRIBUTION: Cuba (Santiago de Cuba).

Scaphioides siboney , new species

Figures 137–146

Stenoonops hoffi (misidentification): Dumitresco and Georgesco, 1983: 94, pl. 16, figs. 1–5.

TYPES: Male holotype, female allotype, plus five male and three female paratypes taken in dry litter at the Reserva Ecológica Siboney-Juticí , 19°57′39″N, 75°42′52″W, Municipio Santiago de Cuba, Santiago de Cuba, Cuba (May 5, 2010; N. Platnick, A. Sánchez, A. Pérez, G. Alayón), deposited in BSC (PBI_OON 964) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. campeche , S. gertschi , and S. hoffi in having sternal pits only around the furrows (figs. 138, 139) and both tubercles and pits with reticulating ridges on the carapace (fig. 137), but have the proximal lobe of the embolus squared in prolateral view (fig. 140) and a widened tip on the anterior genitalic process (figs. 145, 146).

MALE (PBI_OON 964, figs. 137, 138, 140–143): Total length 1.28. Elevated portion of pars cephalica smooth, except posterior margin with short, U-shaped band of tubercles, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture only around furrows, furrows with rows of small pits, anterior margin with interrupted transverse groove. Endite tip with rounded edge. Proximal portion of embolus occupying much more of prolateral side than of retrolateral side.

FEMALE (PBI_OON 964, figs. 139, 144–146): Total length 1.43. Anterior genitalic process very long, greatly expanded anteriorly, base with very short sides.

OTHER MATERIAL EXAMINED: WEST INDIES: Cuba: Santiago de Cuba: Municipio Santiago de Cuba: Cañón del Río Juticí, Reserva Ecológica Siboney-Juticí , 19°57′44″N, 75°44′48″W, May 4–5, 2010, Berlese, litter, elev. 10 m (N. Platnick, A. Sánchez, A. Pérez, G. Alayón, AMNH PBI_OON 965), 2♂ .

DISTRIBUTION: Cuba (Santiago de Cuba).

Scaphioides cobre , new species

Figures 147–159

TYPES: Male holotype, female allotype, and male paratype taken in Berlese sample of litter from El Cobre, 20°02′48″N, 75°56′20″W, Municipio Santiago de Cuba, Santiago de Cuba, Cuba (May 7, 2010; N. Platnick, A. Sánchez, A. Pérez, G. Alayón), deposited in BSC (PBI_OON 966) .

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. bimini , S. camaguey , and S. yateras in having large pits widely distributed on the sternum, including along the midline (figs. 148, 149), but have a scoop-shaped retrolateral margin on the proximal lobe of the embolus (figs. 150, 151) and short, wide anterior genitalic process (figs. 158, 159).

MALE (PBI_OON 966, figs. 147, 148, 150–155): Total length 1.18. Elevated portion of pars cephalica strongly reticulate, posterior margin with U-shaped band of tubercles, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture covering entire surface, furrows with rows of small pits, anterior margin with interrupted transverse groove. Proximal portion of embolus largely covering distal portion in prolateral view.

FEMALE (PBI_OON 966, figs. 149, 156–159): Total length 1.30. Anterior genitalic projection short, wide, expanded anteriorly, situated on semicircular base.

OTHER MATERIAL EXAMINED: None.

DISTRIBUTION: Cuba (Santiago de Cuba).

Scaphioides yateras , new species

Figures 160–167

TYPES: Male holotype and three male paratypes taken 500 m N of the road to Palenque, N of the Río Guayabo , 20°32′37″N, 75°00′41″W, Municipio Yateras, Guantánamo, Cuba (May 13, 2010; A. Pérez), deposited in BSC (PBI_OON 967) ,

ETYMOLOGY: The specific name is a noun in apposition taken from the type locality.

DIAGNOSIS: Members of this species resemble those of S. bimini , S. camaguey , and S. cobre in having large pits widely distributed on the sternum, including along the midline (fig. 161), but have an elongated distal lobe on the embolus (figs. 162, 163).

MALE (PBI_OON 967, figs. 160–167): Total length 1.15. Elevated portion of pars cephalica strongly reticulate, posterior margin with U-shaped band of tubercles, sides strongly reticulate. ALE separated by less than their radius. Sternum surface smooth, covered with large round pits, microsculpture covering entire surface, furrows with rows of small pits, anterior margin with interrupted transverse groove. Distal portion of embolus extending across proximal portion in prolateral view.

FEMALE: Unknown.

OTHER MATERIAL EXAMINED: WEST INDIES: Cuba: Guantánamo: Municipio Baracoa: Río Jaguaní, El Poal , ca. 20°26.050′N, 74°45.775′W, Aug. 11, 1992, elev. 125 m (A. Pérez, M. Estrada, AMNH PBI_OON 43632), 1♂ .

DISTRIBUTION: Cuba (Guantánamo).

Figures

FIGS. 1–15. Scaphioides minuta (Chamberlin and Ivie), male. 1. Habitus, dorsal view. 2. Carapace, lateral view. 3. Same, anterior view. 4. Mouthparts, ventral view. 5. Chelicerae, anterior view. 6. Same, posterior view. 7. Epigastric region, ventral view. 8. Spinnerets, distal view. 9. Anterior lateral spinneret, same. 10. Posterior median spinneret, same. 11. Posterior lateral spinneret, same. 12. Claws of leg I, distal view. 13. Same, leg II. 14. Same, leg III. 15. Same, leg IV.

FIGS. 16–30. Scaphioides minuta (Chamberlin and Ivie), male (16–29) and female (30). 16. Claws of leg I, lateral view. 17. Same, leg II. 18. Same, leg III. 19. Same, leg IV. 20. Tarsal organ from leg I, dorsal view. 21. Same, leg II. 22. Same, leg III. 23. Same, leg IV. 24. Same, palp. 25. Left palp, prolateral view. 26. Same, retrolateral view. 27. Palpal tibia, dorsal view. 28. Left embolus, prolateral view. 29. Same, retrolateral view. 30. Trichobothrial base from tibia I, dorsal view.

FIGS. 31–45. Scaphioides minuta (Chamberlin and Ivie), female. 31. Habitus, dorsal view. 32. Same, lateral view. 33. Carapace, lateral view. 34. Same, anterior view. 35. Mouthparts, ventral view. 36. Chelicerae, anterior view. 37. Same, posterior view. 38. Endite and labrum, dorsal view. 39. Serrula, dorsal view. 40. Palp, prolateral view. 41. Same, retrolateral view. 42. Palpal tibia, dorsal view. 43. Epigastric region, ventral view. 44. Genitalia, dorsal view. 45. Spinnerets, distal view.

FIGS. 46–60. Scaphioides minuta (Chamberlin and Ivie), female. 46. Anterior lateral spinneret, distal view. 47. Posterior median spinneret, same. 48. Posterior lateral spinneret, same. 49. Claws of leg I, distal view. 50. Same, leg II. 51. Same, leg III. 52. Same, leg IV. 53. Claws of leg I, lateral view. 54. Same, leg III. 55. Same, leg IV. 56. Tarsal organ from leg I, dorsal view. 57. Same, leg II. 58. Same, leg III. 59. Same, leg IV. 60. Same, palp.

FIGS. 61–68. Scaphioides minuta (Chamberlin and Ivie), male (61–64) and female (65–68). 61, 65. Carapace, dorsal view. 62, 66. Sternum, ventral view. 63. Left embolus, prolateral view. 64. Same, retrolateral view. 67. Genitalia, ventral view. 68. Same, dorsal view.

FIGS. 69–78. Scaphioides minuta (Chamberlin and Ivie), male (69–73) and female (74), S. bimini, new species, male (75–78). 69, 75. Carapace, dorsal view. 70. Left palp, prolateral view. 71. Same, retrolateral view. 72, 77. Left embolus, prolateral view. 73, 78. Same, retrolateral view. 74. Epigastric region, ventral view. 76. Sternum, ventral view.

FIGS. 92–100. Scaphioides irazu, new species, female. 92, 96. Carapace, dorsal view. 93. Sternum, ventral view. 94. Habitus, dorsal view. 95. Same, ventral view. 97. Carapace, lateral view. 98. Epigastric region, ventral view. 99. Genitalia, ventral view. 100. Same, dorsal view.

FIGS. 101–113. Scaphioides gertschi, new species, male (101, 102, 104–108, 110, 111) and female (103, 109, 112, 113). 101, 106. Carapace, dorsal view. 102, 103. Sternum, ventral view. 104, 110. Left embolus, prolateral view. 105, 111. Same, retrolateral view. 107. Left palp, prolateral view. 108. Same, retrolateral view. 109. Epigastric region, ventral view. 112. Genitalia, ventral view. 113. Same, dorsal view.

FIGS. 114–126. Scaphioides camaguey, new species, male (114, 115, 117–122) and female (116, 123–126). 114, 119, 123. Carapace, dorsal view. 115, 116. Sternum, ventral view. 117. Left embolus, prolateral view. 118. Same, retrolateral view. 120. Left palp, prolateral view. 121. Same, retrolateral view. 122. Mouthparts, ventral view. 124. Epigastric region, ventral view. 125. Genitalia, ventral view. 126. Same, dorsal view.

FIGS. 127–136. Scaphioides granpiedra, new species, male (127, 128, 130–133) and female (129, 134–136). 127. Carapace, dorsal view. 128, 129. Sternum, ventral view. 130. Left embolus, prolateral view. 131. Same, retrolateral view. 132. Left palp, prolateral view. 133. Same, retrolateral view. 134. Epigastric region, ventral view. 135. Genitalia, ventral view. 136. Same, dorsal view.

FIGS. 137–146. Scaphioides siboney, new species, male (137, 138, 140–143) and female (139, 144–146). 137. Carapace, dorsal view. 138, 139. Sternum, ventral view. 140. Left embolus, prolateral view. 141. Same, retrolateral view. 142. Left palp, prolateral view. 143. Same, retrolateral view. 144. Epigastric region, ventral view. 145. Genitalia, ventral view. 146. Same, dorsal view.

FIGS. 160–167. Scaphioides yateras, new species, male. 160, 164. Carapace, dorsal view. 161. Sternum, ventral view. 162. Left embolus, prolateral view. 163. Same, retrolateral view. 165. Mouthparts, ventral view. 166. Left palp, prolateral view. 167. Same, retrolateral view.