Haplodrassus lyndae Abrous & Bosmans, 2018

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67 : 17-22

Haplodrassus lyndae Abrous & Bosmans , sp. n.

Figs 58‒66 , 77‒82 , map 2

Types: Holotype ♂, paratypes 4 ♂ and 4♀ from Algeria, wilaya Djelfa, Djébel Sénalba (N34°33'50” E3°2'9”), 1310‒1400m, pitfalls in Pinus halepensis forest, 1990–1991, N. Bouragba & R. Bosmans leg.; holotype, 2♂ and 2♀ paratypes deposited in RBINS, 2♂ 2♀ paratypes deposited in MNHN ( MNHN AR15696 ) .

MAP 2. Distribution of Haplodrassus dentifer sp. n. (circles) and H. lyndae sp. n. (squares). MAP 3. Distribution of Haplodrassus longivulva sp. n. (squares), H. ovatus sp. n. (circles) and H. rhodanicus sp. n. (triangles).

Diagnosis. This species is easily distinguished from other Haplodrassus species of the Maghreb, in the male by the 2 blunt, widely separated terminal teeth of the embolic apophysis ( Figs 62 , 79 ), in the female by the narrow fovea, the hood being 3 times as wide as the fovea ( Figs 64 , 81 ).

Etymology. The species is dedicated to our friend Lynda Beladjal, in recognition of her precious aid during this study.

Description. Measurements: Male (n=10): total length 2.9‒5.7; carapace 1.4‒2.7 long, 0.9‒1.6 wide. Female (n=10): total length 3.7‒6.4; carapace 1.7‒2.7 long, 1.3‒1.9 wide. Colour: Carapace yellowish brown, cephalic part slightly darkened; legs yellowish brown; abdomen pale grey, rarely with indications of whitish chevrons. Spination: General pattern.

Male palp ( Figs 60‒63 , 77‒80 ): Tibial apophysis as long as the tibia’s diameter, gradually narrowing, terminally widened and asymmetric; embolic apophysis with parallel margins, slightly bent, terminally with two blunt teeth.

Epigyne ( Figs 64‒66 , 81‒82 ): Longer than wide (ratio 1:1.5), pro-fovea very narrow; fovea elongated, narrow anteriorly, dilated in the posterior half; lateral pockets oval, with small lateral pouch.

Further material examined. AFRICA, ALGERIA, Batna: Forêt de S'Gag (N35°26’1” E6°14’38”), 1650m, 2♂, pitfalls in Cedrus atlantica forest, 5.XI.1987 ‒ 9.IV.1988, Massif de l’Aurès, Belezma, Col Telmet (N35°35'11” E6°2'50”), 1800m, 3♂ 8♀, pitfalls in Cedrus atlantica forest, 28.II.1988 (CRB), Blida: Atlas Blidéen, Chréa, les Glacières (N36°26’44” E2°56’22”), 1045m, 1♂, pitfalls in dense Quercus ilex forest, 20.X.1987, idem, 1520m, Pic Abdelkader (N36°24’14” E2°50’6”), 1♀, pitfalls in Cedrus atlantica forest, 20.VI.1987 (CRB), Chréa, south slope (N36°23’38” E2°48’24”), 3♂, pitfalls in dense Cedrus atlantica forest, 15.II.1987 ‒ 21.VI.1987 (COA), Djelfa: Djébel Sénalba (N34°33'50” E3°2'9”), 1220‒1450m, 2♂ 8♀, pitfalls in Pinus halepensis forest, 1990‒1991, (CNB, CRB), idem, 1310‒1400m, 3♂ 4♀, 1990‒1991 (CNB, CRB), M’sila: Aïn Oghrab (N35°0’52” E4°6’24”), 650m, 1♀, pitfalls in Pinus halepensis forest, 20.III‒23.IV.1990 (CRB), Tiaret: between Rosfa and Aïn el Hadid (N34°58'59” E0°48'52”), 900m, 1♂, pitfalls in dry Pinus halepensis forest, 22.V.1990 (CRB). MOROCCO, Draâ-Tafilalet, Bouldjoul N., Col du Zad (N33°0'36” W5°3"36”), 2280m, 1♀, under stones in degraded Cedrus atlantica and Quercus ilex forest, 7.II.1996, J. Van Keer leg. (CJVK), Marrakech-Safi, Marrakech N., Oued Tensift (N31°41'33” W7°59'17”), 370m, 1♀, stones in dry river bed, 9.II.1996, J. Van Keer leg. (CJVK), Rabat-Salé-Kénitra, Forêt de Maâmora, Sidi Allal-Bahraoui (N34°2'41” W6°35'25”), 175m, 1♂ 2 subadult♀, 2.X.1989, 1♂, 6.XI.1989, 2♀, 5.XII.1989, litter in Quercus suber forest, S. Benhalima leg. (CSB), idem, 2♀, 8.II.1996, R. Bosmans leg. (CRB), Tanger-Tetouan-Al Hoceima, Issaguen S. (N34°53'25” W4°34'59”), 1450m, 1♀, stones in Cedrus atlantica forest, 15.XII.2013 (CRB). EUROPE, SPAIN, Cuenca: Molinia de Aragon (N40°51’39” W1°49’58), 1050m, 1♀, stones in Pinus forest, 14.IV.1998 (CRB), Jaen: Jaen S., Jabalcuz (N37°43'52” W3°48'33”), 700m, 1♀, stones in Pinus forest, 12.IV.1998 (CRB), Madrid: El Escorial (N40°35’29” W4°08’51”), 1♀, as Drassodes virens L. Koch nomen nudum, Coll. Simon 2049 (MNHN AR9375).

Remark. The following material possibly belongs to H. lyndae sp. n., but the capture of males should confirm this: ITALY, Sicilia, Terrasini, Riserva Naturale Orientata Capo Rama (N38°8'22” E13°3'38”), 2♀, pitfalls in grassland, 14‒ 24.IV.2012, C. Cusimano leg. (MCSNB).

Ecology. The species is common in the Cedrus and Quercus forests at higher altitudes of Algeria and Morocco. Males were collected from November to May, females from December to June.

Distribution. Central and North Morocco, North Algeria, Central and South Spain and possibly Italy (see above; map 2). It is remarkable the species is present in some Algerian stations of high altitudes (Atlas Blidéen, Aurès Massif, Djébel Sénalba), but absent from others (Massif du Djurdjura, Djébel Ouarsenis and Djébel Babor), where also collections were made at regular intervals.

Figures

FIGURES 58–66. Haplodrassus lyndae sp. n. (Algeria, Col Telmet). 58. Male, dorsal aspect; 59. Female, dorsal aspect; 60. Male palpal tibia, dorsolateral view; 61. Male left palp, retrolateral view; 62. Idem, ventral view; 63. Idem, prolateral view; 64. Epigyne, ventral view; 65. Vulva, ventral view; 66. Vulva, dorsal view.

FIGURES 77–86. Figs 77‒82. Haplodrassus lyndae sp. n. (Algeria, Col Telmet). 77. Male left palp, ventral view; 78. Male palpal tibia, dorsolateral view; 79. Embolic apophysis of male palp, ventral view; 80. Embolus, dorsal view; 81. Epigyne, ventral view; 82. Vulva, ventral view. Figs 83‒86. Haplodrassus nigroscriptus (type series from Morocco, Essaouira). 83. Male left palp, ventral view; 84. Male palpal tibia, dorsolateral view; 85. Epigyne, ventral view; 86. Vulva, ventral view.