Haplodrassus, Chamberlin, 1922

Bosmans, Robert, Kherbouche-Abrous, Ourida, Benhalima, Souâd & Hervé, Christophe, 2018, The genus Haplodrassus Chamberlin, 1922 in the Mediterranean and the Maghreb in particular (Araneae: Gnaphosidae), Zootaxa 4451 (1), pp. 1-67: 7-8

publication ID

https://doi.org/10.11646/zootaxa.4451.1.1

publication LSID

lsid:zoobank.org:pub:5DCE18B3-9DBA-48F5-AEF7-90A50B984EA5

persistent identifier

http://treatment.plazi.org/id/DE5087A6-C013-B331-FF39-7404FA53FC3E

treatment provided by

Plazi

scientific name

Haplodrassus
status

 

Key to the Haplodrassus   species of the Maghreb region

MALES

1 Cymbium with abundant, strong spines (fig. 71)................................................... nigroscriptus  

- Cymbium with few, less strong spines (fig. 32).............................................................. 2

2(1) Embolic apophysis hooked (fig. 190)........................................................ triangularis   sp. n.

2 Embolic apophysis sigmoid or nearly straight ( Figs 4 View FIGURES 1–9 , 14 View FIGURES 10–17 , 35 View FIGURES 30–40 …)................................................. 3

3(2) Embolic apophysis small, occupying only one third of tegulum ( Figs 122 View FIGURES 118–125 , 136 View FIGURES 136–144 ); tibial apophysis slender, with narrow base ( Figs 120 View FIGURES 118–125 , 137 View FIGURES 136–144 )............................................................................... rhodanicus  

- Embolic apophysis large, occupying at least half of tegulum ( Figs 4 View FIGURES 1–9 , 14 View FIGURES 10–17 , 35 View FIGURES 30–40 …); tibial apophysis voluminous, with broad base ( Figs 5 View FIGURES 1–9 , 12 View FIGURES 10–17 , 33 View FIGURES 30–40 …)..................................................................................... 4

4(3) Embolic apophysis with dorsal keel ( Figs 158 View FIGURES 154–161 , 169 View FIGURES 162–171 )..................................................... signifer  

- Embolic apophysis without dorsal keel ( Figs 4 View FIGURES 1–9 , 14 View FIGURES 10–17 , 35 View FIGURES 30–40 …)...................................................... 5

5(4) Embolic apophysis with 2 prolateral teeth ( Figs 14 View FIGURES 10–17 , 26 View FIGURES 18–29 , 93 View FIGURES 87–97 , 110 View FIGURES 108–117 )................................................. 6

- Embolic apophysis without ( Figs 4 View FIGURES 1–9 , 35 View FIGURES 30–40 , 103 View FIGURES 98–107 …) or at most with 1 prolateral tooth ( Figs 62 View FIGURES 58–66 , 129 View FIGURES 126–135 )....................... 7

6(5) Embolic apophysis slender, elongated and with pointed tip ( Figs 93 View FIGURES 87–97 , 108 View FIGURES 108–117 ).................................... omissus  

- Embolic apophysis compact, robust, with blunt tip ( Figs 14 View FIGURES 10–17 , 25 View FIGURES 18–29 )........................................ dalmatensis  

7(6) Embolic apophysis with one prolateral tooth ( Figs 62 View FIGURES 58–66 , 77 View FIGURES 77–86 , 129 View FIGURES 126–135 , 140 View FIGURES 136–144 )............................................. 8

- Embolic apophysis without prolateral tooth ( Figs 4 View FIGURES 1–9 , 35 View FIGURES 30–40 , 103 View FIGURES 98–107 …)................................................. 9

8(7) Teeth on embolic apophysis equal, apophysis appearing bifid ( Figs 62 View FIGURES 58–66 , 79 View FIGURES 77–86 )............................... lyndae   sp. n.

- Teeth on embolic apophysis not equal (fig. 129, 142).................................................... rufipes  

9(8) Embolus with a strong retrolateral tooth at its base ( Figs 49, 51 View FIGURES 49–57 )....................................... dentifer   sp. n.

- Embolus without retrolateral tooth at base ( Figs 4 View FIGURES 1–9 , 45 View FIGURES 41–48 , 103 View FIGURES 98–107 …)............................................ …….10

10(9) Tibial apophysis nearly symmetric ( Figs 100 View FIGURES 98–107 , 114 View FIGURES 108–117 ).................................................. ovatus   sp. n.

- Tibial apophysis strongly asymmetric ( Figs 5 View FIGURES 1–9 , 43 View FIGURES 41–48 , 147 View FIGURES 145–153 , 186 View FIGURES 181–189 )................................................... 11

11(10) Tibial apophysis terminally strongly pointed ( Figs 186 View FIGURES 181–189 , 195 View FIGURES 190–197 ).............................................. typhon  

- Tibial apophysis bluntly pointed ( Figs 5 View FIGURES 1–9 , 43 View FIGURES 41–48 , 147 View FIGURES 145–153 )........................................................... 12

12(11) Tibial apophysis gradually narrowing, with subterminal concavity ( Figs 43 View FIGURES 41–48 , 55 View FIGURES 49–57 )....................... longivulva   sp. n.

- Tibial apophysis not gradually narrowing, with basal concavity ( Figs 5 View FIGURES 1–9 , 147 View FIGURES 145–153 )..................................... 13

13(12) Embolic apophysis terminally with pointed nipple ( Figs 6 View FIGURES 1–9 , 20 View FIGURES 18–29 )........................................... crassipes  

- Embolic apophysis terminally blunt ( Figs 149 View FIGURES 145–153 , 164 View FIGURES 162–171 )............................................... securifer   sp. n. FEMALES

1 Tarsus and tibia of palp with more than 20 ventral spines (fig. 73)..................................... nigroscriptus  

- Tarsus of palp with less than 10 ventral spines (fig. 37........................................................ 2

2(1) Fovea triangular ( Figs 178 View FIGURES 172–180 , 192 View FIGURES 190–197 )............................................................ triangularis   sp. n.

- Fovea oval or rounded ( Figs 7 View FIGURES 1–9 , 16 View FIGURES 10–17 , 38 View FIGURES 30–40 …)................................................................... 3

3(2) Epigyne at least twice as long as wide ( Figs 47 View FIGURES 41–48 , 64 View FIGURES 58–66 , 124 View FIGURES 118–125 )....................................................... 4

- Epigyne less elongate, at most 1.7 times as long as wide ( Figs 7 View FIGURES 1–9 , 16 View FIGURES 10–17 , 38 View FIGURES 30–40 ).......................................... 6

4(3) Fovea narrow anteriorly, widening distinctly to the back ( Figs 64 View FIGURES 58–66 , 81 View FIGURES 77–86 )................................... lyndae   sp. n.

- Fovea more or less rectangular ( Figs 47 View FIGURES 41–48 , 124 View FIGURES 118–125 )............................................................... 5

5(4) Lateral margin of fovea with median angularity ( Figs 47 View FIGURES 41–48 , 56 View FIGURES 49–57 )...................................... longivulva   sp. n.

- Lateral margin of fovea with parallel margins ( Figs 124 View FIGURES 118–125 , 138 View FIGURES 136–144 )........................................... rhodanicus  

6(5) Fovea with postero-median septum ( Figs 16 View FIGURES 10–17 , 95 View FIGURES 87–97 )............................................................. 7

- Fovea without postero-median septum ( Figs 7 View FIGURES 1–9 , 38 View FIGURES 30–40 , 105 View FIGURES 98–107 )....................................................... 8

7(6) Fovea distinctly longer than wide ( Figs 17 View FIGURES 10–17 , 28 View FIGURES 18–29 )..................................................... dalmatensis  

- Fovea as long as wide ( Figs 95 View FIGURES 87–97 , 111 View FIGURES 108–117 )................................................................. omissus  

8(7) Foveal margin with distinct median tooth in lateral margin, dividing fovea into a trapezoid or rectangular anterior part and a rounded posterior part ( Figs 40 View FIGURES 30–40 , 132 View FIGURES 126–135 )...................................................................... 9

- Foveal margin without such tooth, fovea oval or rounded ( Figs 7 View FIGURES 1–9 , 105 View FIGURES 98–107 , 151 View FIGURES 145–153 , 160 View FIGURES 154–161 , 187 View FIGURES 181–189 ).............................. 10

9(8) Areola anteriorly truncate ( Figs 132 View FIGURES 126–135 , 143 View FIGURES 136–144 )............................................................. rufipes  

- Areola anteriorly gradually narrowing ( Figs 40 View FIGURES 30–40 , 52 View FIGURES 49–57 )................................................ dentifer   sp. n.

10(9) Fovea longer than wide ( Figs 105 View FIGURES 98–107 , 187 View FIGURES 181–189 )................................................................... 11

- Fovea as long as wide ( Figs 7 View FIGURES 1–9 , 151 View FIGURES 145–153 , 160 View FIGURES 154–161 ).................................................................. 12

11(10) Fovea oval, anterior margins converging ( Figs 105 View FIGURES 98–107 , 116 View FIGURES 108–117 ).............................................. ovatus   sp. n.

- Fovea not oval, anterior margins diverging ( Figs 187 View FIGURES 181–189 , 196 View FIGURES 190–197 )................................................ typhon  

12(11) Areola elongated, anteriorly continuing into a chitinised crest ( Figs 160 View FIGURES 154–161 , 170 View FIGURES 162–171 )................................. signifer  

- Areola less elongated, anteriorly rounded ( Figs 7 View FIGURES 1–9 , 151 View FIGURES 145–153 )....................................................... 13

13(12) Epigyne as long as wide ( Figs 151 View FIGURES 145–153 , 165 View FIGURES 162–171 ), lateral pockets nearly rounded ( Figs 152 View FIGURES 145–153 , 166 View FIGURES 162–171 )................... securifer   sp. n.

- Epigyne longer than wide, ( Figs 7 View FIGURES 1–9 , 22 View FIGURES 18–29 ), lateral pockets more oval ( Figs 9 View FIGURES 1–9 , 23 View FIGURES 18–29 )............................... crassipes