Eusphalerum sp. 2.

Shavrin, Alexey V. & Yamamoto, Shuhei, 2019, Unexpected palaeodiversity of omaliine rove beetles in Eocene Baltic amber (Coleoptera, Staphylinidae, Omaliinae), ZooKeys 863, pp. 35-83 : 35

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Eusphalerum sp. 2.


Eusphalerum sp. 2. Figures 12, 13, 81-82, 83-85

Materials examined.

One male, complete specimen as an inclusion in a piece of yellow Baltic amber 35.4 mm × 21.5 mm × 7.5 mm in size (Figs 12, 13), with glued very small quadrate paper on plastic bag labeled "AWI | 045", with the following labels: "AWI-045 | Phyllodrepa (?) | 3 spec." <rectangular handwritten label>, "Dr. Vitalii Alekseev's | Collection" <rectangular handwritten label>, " Eusphalerum sp. 2 | Shavrin A.V. det. 2018" <rectangular label, printed>. The specimen is deposited in the private collection of Vitalii I. Alekseev (Kaliningrad, Russia), registered as AWI-045.


The single specimen is a male located close to the outer surface of the piece of amber, with many details visible in both dorsal and ventral surfaces. The elytra are somewhat deformed and seem flattened, and the right elytron is depressed into the thorax. Additionally, the piece of the amber contains two males of Eu. sp. 3 and Eu. sp. 4 (see below), and a syninclusion located near the narrowest side of the amber: nymph of small mite about 0.50 mm in length (Figs 12, 13).


Measurements: HW: 0.53; HL: 0.20; OL: 0.11; AL: 0.69; PML × PMW (III, IV): III: 0.03 × 0.01, IV: 0.07 × 0.02; PL: 0.43; PW: 0.67; ESL: 0.85; EW: 0.81; MTbL: 0.36; MTrL: 0.27 ( I–IV: 0.14; V: 0.13); AW: 0.79; TL: 2.06. Antennomeres with lengths × widths: 1: 0.12 × 0.02; 2: 0.07 × 0.02; 3: 0.06 × 0.02; 4: 0.05 × 0.02; 5-6: 0.05 × 0.03; 7: 0.04 × 0.04; 8: 0.05 × 0.04; 9-10: 0.05 × 0.05; 11: 0.10 × 0.04.

Body elongate, somewhat flattened (Fig. 81), glossy and glabrous, without visible setation. Body appears dark-brown, with basal portions of pronotum and legs reddish-brown. Body dorsolaterally as in Figure 82 and ventrally as in Figure 83.

Head strongly transverse, distinctly more than twice as wide as long, with slightly convex middle portion and posterior parts of infraorbital ridges, without visible grooves in front of ocelli and postocular carina (Fig. 84). Head with indistinct, small and sparse punctation, with moderately coarse and large isodiametric microsculpture between punctures becoming more transverse toward middle part of neck. Eyes large, strongly protruding laterad. Ocelli small, convex, situated very close to infraorbital ridges at level of about middle length of eyes, distance between ocelli about twice as long as distance between ocellus and margin of eye (Figs 82, 84). Labrum transverse. Apical segment of maxillary palp elongate, slightly wider in middle than penultimate segment, from apical third gradually narrowed toward moderately acute apex (Figs 82, 84). Gular sutures not fully visible, widely separated from each other (Fig. 83). Antenna moderately long, distinctly exceeding shoulders of elytra, with elongate setae, distinctly longer on antennomeres 6-11; basal antennomere wide, gradually widened apicad, antennomere 2 similar in width, swollen and elongate, 3 with thin basal portion, slightly widened apicad, 4 slightly shorter than 3, 5, and 6 slightly wider than 4, 7 short and moderately rounded, 8-10 slightly transverse, apical antennomere more than twice as long as broad, from about middle strongly narrowed toward acute apex (Figs 81-84).

Pronotum 1.5 times as wide as long, slightly broader than head, widest in middle, markedly more narrowed posterad than anterad; apical margin slightly and widely rounded, about as broad as posterior margin, anterior and posterior angles widely rounded; laterobasal margins slightly concaved, with very indistinct small crenulation; lateral margins narrowly explanate; lateral portions with indistinct semioval impression about middle (Figs 81, 84). Pronotum with somewhat regular small and sparse punctation and with isodiametric ground sculpture slightly coarser than that on head (Figs 81, 84). Prosternum with moderately wide and protruded prosternal process (Fig. 83). Scutellum large and wide (Fig. 81).

Elytra little longer than wide, about twice as long as pronotum, gradually widened apicad, reaching basal to apical margins of abdominal tergite IV, with widely rounded apicolateral angles; shoulders moderately widely rounded; lateral edges narrowly explanate (Fig. 81). Punctation of elytra invisible in details but appears slightly denser and deeper than that on pronotum.

Legs moderately long and slender, femora markedly widened in middle, tibiae moderately short and thin, gradually widened apicad, slightly shorter than femora, covered by elongate setae, with a few strong setae on apical margins near apex; tarsomeres 1-4 distinctly wide, with dense and long setae; apical metatarsomere long, yet slightly shorter than length of preceding tarsomeres together; tarsal claws simple, elongate (Figs 81-83).

Abdomen (Fig. 85) slightly narrower than elytra; abdominal tergites with sparse and moderately small punctures, no wing-folding patches are visible.

Male. Apical margin of abdominal tergite VIII slightly rounded. Apical margin of abdominal sternite VIII slightly sinuate.

Female unknown.


As in the previous species, this specimen has very long and moderately dense setae on lateral portions of tarsomeres 1-4, distinctly deformed body (especially elytra) and unusually strongly protruded eyes.