Dasyhelea aliciae, Grogan & Díaz & Spinelli & Ronderos, 2019
publication ID |
https://doi.org/ 10.11646/zootaxa.4700.3.1 |
publication LSID |
lsid:zoobank.org:pub:B0712990-7A3B-4042-9A4B-5CE96AFAECF8 |
DOI |
https://doi.org/10.5281/zenodo.5457122 |
persistent identifier |
https://treatment.plazi.org/id/DF72B52D-DE04-FF8A-FF54-1CEBFA1B0897 |
treatment provided by |
Plazi |
scientific name |
Dasyhelea aliciae |
status |
sp. nov. |
Key to adult Dasyhelea View in CoL View at ENA of Curaçao (females unknown for D. aliciae n. sp. & D. recurva n. sp.)
1 Males .............................................................................................. 2
- Females............................................................................................ 13
2 Aedeagus nearly covered with, or, only distal portion covered with thin hyaline envelope ( Fig. 69 View FIGURES 52–69 ); gonocoxal apodemes and paramere symmetrical ( Fig. 68 View FIGURES 52–69 )......................................................................... 3
- Aedeagus not covered with thin hyaline envelope ( Figs. 54, 57, 60, 63, 66 View FIGURES 52–69 ); gonocoxal apodemes and paramere usually asymmetrical, rarely symmetrical ( Figs. 53, 56, 59, 62, 65 View FIGURES 52–69 )........................................................ 5
3 Small species; wing length 0.85 mm, wing very broad, membrane without macrotrichia; paramere straight, club-shaped; aedeagus very complex, composed of two sections, a heavily sclerotized H-shaped basal portion and a broad, hyaline membranous quadrate posterior portion attached to the distal section of the basal portion, the apex of which is broad with short, blunt apicolateral extensions and an apicocentral lobe ( Figs. 45 View FIGURES 42–51 , 67–69 View FIGURES 52–69 ) (unplaced to species group)... D. rhopaloparamera View in CoL n. sp.
- Large species; wing length 1.25–1.50 mm, wing narrow, membrane with long macrotrichia, densest apically; paramere straight, tapered distally to pointed or rounded tip; aedeagus nearly entirely covered with thin hyaline envelope (cincta group)...... 4
4 Paramere elongate, slender with sharply pointed tip and base fused to both gonocoxal apodemes; bases of gonocoxal apodemes broad, foot-shaped; distal margin of hyaline envelope of aedeagus without crenulations.............. D. cincta (Coquillett) View in CoL
- Paramere relatively short, rod-like and not usually fused to gonocoxal apodemes; bases of gonocoxal apodemes slender, recurved; distal margin of hyaline envelope of aedeagus with crenulations...................... D. bahamensis (Johnson) View in CoL
5 Frontal sclerite broad, elliptical, bilobate or heart-shaped; antennal flagellomeres 2–13 heavily sculptured, 13 elongate with a tapered slender apex; gonocoxal apodemes and paramere asymmetrical; gonocoxite often with basomesal hook or spur (grisea group)............................................................................................. 6
- Frontal sclerite slender, elliptical with or without a pair of lower marginal ear-like lobes; antennal flagellomeres 2–12 usually not sculptured, 13 short to moderately long with rounded apex; gonocoxal apodemes and paramere usually asymmetrical, rarely symmetrical; gonocoxite without basomesal hook or spur..................................................... 8
6 Aedeagus with unequal length posterolateral arms, right arm shorter than left arm; sternite 9 mid-portion margin with numerous small spicules................................................................... D. azteca Huerta & Grogan View in CoL
- Aedeagus with equal length posterolateral arms; sternite 9 mid-portion margin without numerous small spicules......... 7
7 Gonocoxite with large basomesal hook; paramere fused to both gonocoxal apodemes, apex curved or straight, not bent ventrad................................................................................ D. grisea (Coquillett) View in CoL
- Gonocoxite with or without small basomesal spur; paramere sometimes fused to only base of right gonocoxal apodeme, apex bent ventrad................................................................ D. flavifrons (Guérin-Méneville) View in CoL
8 Sternite 9 distal margin extending to just below basal arch or as far as beyond apex of aedeagus ( Fig. 5 View FIGURES 1–5 ); frontal sclerite usually with pair of lower marginal ear-like lobes; aedeagus usually without a posteromedian projection ( Fig. 54 View FIGURES 52–69 ); distal margin of tergite 9 usually with short conical apicolateral processes (leptobranchia group)................................... 9
- Sternite 9 distal margin concave, straight or extending to base of or onto aedeagus but not beyond apex (Figs. 10, 15, 25, 34); frontal sclerite elliptical, without lower marginal ear-like lobes; aedeagus with a posteromedian projection ( Figs. 57, 60, 63, 66 View FIGURES 52–69 ); distal margin of tergite 9 usually with elongate conical or cylindrical apicolateral processes (mutabilis group)....... 10
9 Sternite 9 mid-portion greatly elongate, extending beyond apex of aedeagus, distal section expanded laterally; paramere relatively short, distal portion not recurved ventrally; tergite 9 apicolateral processes parallel.......... D. corinneae Gosseries View in CoL
- Sternite 9 mid-portion moderately elongate, extending just below basal arch of aedeagus, distal margin slightly curved; paramere long, distal portion elongate, slender, recurved ventrally; tergite 9 apicolateral processes divergent ( Figs. 5 View FIGURES 1–5 , 52–53 View FIGURES 52–69 )............................................................................................. D. aliciae View in CoL n. sp.
10 Paramere without tapered, recurved apical section ( Fig. 59 View FIGURES 52–69 ).................................................. 11
- Paramere with tapered, recurved apical section ( Figs. 62, 65 View FIGURES 52–69 )................................................. 12
11 Paramere asymmetrical, broad basally with angled apex; aedeagus broad, heavily sclerotized, apical portion bifurcate with bifid, recurved apices; tergite 9 with parallel apicolateral processes (Figs. 10, 56–57)................... D. recurva View in CoL n. sp.
- Paramere nearly symmetrical, slender basally with bifid apex; aedeagus complex, composed of two sections, a heavily sclerotized basal portion and a hyaline distal portion; tergite 9 with broadly divergent apicolateral processes ( Figs. 15 View FIGURES 11–21 , 58–60 View FIGURES 52–69 )............................................................................................ D. latiala View in CoL n. sp.
12 Gonostylus elongate with very slender, recurved apical section; gonocoxal apodemes broadly fused with paramere; apical section of paramere abruptly recurved, adpressed to ventral side of paramere; aedeagus with very shallow basal arch, posterolateral arms with mesal short, slender, subapical toothlike process ( Figs. 25 View FIGURES 22–30 , 62–63 View FIGURES 52–69 )........................ D. cyrtostyla View in CoL n. sp.
- Gonostylus moderately short with slender rounded apex; only right gonocoxal apodeme fused with paramere; apical section of paramere recurved, J-shaped, not adpressed to paramere; aedeagus with deep, curved basal arch, posterolateral arms with pointed apices, without mesal subapical toothlike process ( Figs. 34 View FIGURES 31–41 , 65–66 View FIGURES 52–69 )...................... D. curacaoensis View in CoL n. sp.
13 Two spermathecae (cincta group)....................................................................... 14
- One spermatheca.................................................................................... 15
14 Antennal flagellomeres 9–13 elongate, much longer than broad….. D. cincta (Coquillett) View in CoL
- Antennal flagellomeres 9–12 very short, only 13 slightly elongate, longer than broad............ D. bahamensis (Johnson) View in CoL
15 Frontal sclerite broad, elliptical, bilobate or heart-shaped; antennal flagellomeres 2–13 usually heavily sculptured, 13 elongate with a tapered slender apex; subgenital plate usually with triangular, arrowhead-shaped or slender distal portion (grisea group)............................................................................................ 16
- Frontal sclerite slender, elliptical, with or without lower marginal ear-like lobes; antennal flagellomeres usually not heavily sculptured, flagellomere 13 short to moderately long with rounded apex; subgenital plate circular with a central lumen, disjunct, or, slender bridge-like................................................................................. 18
16 Subgenital plate with a pair of basolateral pointed projections and distal portion arrowhead-shaped; spermatheca subspherical, often retort-shaped with a long neck that is usually recurved.......................... D. flavifrons (Guérin-Méneville) View in CoL
- Subgenital plate without basolateral pointed projections, distal portion arrowhead-shaped or slender; spermatheca pyriform, with or without long slender neck....................................................................... 17
17 Subgenital plate lightly to moderately sclerotized, distal portion slender or arrowhead-shaped; spermatheca pyriform with large well-developed surface punctations, distal portion curved with long slender neck; flagellomeres 9-12 without sculpturing, only 13 with distinct sculpturing........................................................ D. azteca Huerta & Grogan View in CoL
- Subgenital plate heavily sclerotized with distinct arrowhead-shaped distal portion; spermatheca pyriform without numerous large, well-developed surface punctations, without a long neck; flagellomeres 9-13 with distinct sculpturing....... D. grisea (Coquillett) View in CoL
18 Subgenital plate slender, bridge-like, without central lumen, or, arched at mid-length with poorly developed central lumen ( Fig. 51 View FIGURES 42–51 ); antennal flagellomeres 3–13 elongate or only 13 elongate ( Fig. 46 View FIGURES 42–51 )......................................... 19
- Subgenital plate circular or ovoidal with central lumen ( Figs. 21 View FIGURES 11–21 , 30 View FIGURES 22–30 , 41 View FIGURES 31–41 ); antennal flagellomeres 3–12 usually relatively short, 9–13 sometimes longer than broad ( Figs. 26 View FIGURES 22–30 , 35 View FIGURES 31–41 ) (mutabilis group)............................................. 20
19 Antennal flagellomeres 2–12 vasiform, slightly longer than broad, only 13 elongate, much longer than broad; subgenital plate slender, abruptly elevated at mid length with a narrow disjunct curved sclerite and quadrate central lumen; wing membrane nearly devoid of macrotrichia, a few in margins of cells r 3 and m 1; spermatheca spheroidal, large (67 x 60 µm) with long, slender neck; sternite 8 with 2 separated lengthwise groups of 3–6 long setae ( Figs. 46, 50–51 View FIGURES 42–51 ) (unplaced to species group)...................................................................................... D. rhopaloparamera View in CoL n. sp.
- Antennal flagellomeres 2–13 elongate, 9-13 much longer than broad; subgenital plate slender, bridge-like, without disjunct anterior sclerite; wing membrane with dense macrotrichia in all cells; sternite 8 with mid lateral row of 4–8 long setae; spermatheca ovoid, small (45 x 30 µm) with short slender neck (leptobranchia group)…………. D. corinneae Gosseries View in CoL
- 20 Antennal flagellum very short (total length 0.31-0.33 mm), flagellomeres 2–12 very short, moniliform, only 13 longer than broad; wing with very short costa, costal ratio 0.23–25; subgenital plate with small narrow lumen; spermatheca small (35 x 28 µm) ( Figs. 35, 40–41 View FIGURES 31–41 )................................................................ D. curacaoensis View in CoL n. sp.
- Antennal flagellum elongate to moderately long (total length 0.36–0.48 mm), flagellomeres 9–13 slightly to moderately elongate, longer than 2–8; wing with moderately long costa, costal ratio 0.44–0.50; subgenital plate with large broad lumen; spermatheca medium-size to large (42.5 x 37.5 µm–57.5 x 32.5 µm)............................................... 21
21 Antennal flagellomeres 9–13 elongate, much longer than 2–8; wing very broad with moderately long costa, costal ratio 0.50; subgenital plate with a sub-circular basal portion with a large ovoid lumen and a very broad trapezoidal apical portion; spermatheca medium-size (42.5 x 37.5 µm), ovoid, heavily sclerotized, neck short, slightly recurved ( Figs. 16, 20–21 View FIGURES 11–21 )................................................................................................... D. latiala View in CoL n. sp.
- Antennal flagellomeres 2–12 slightly longer than broad, only 13 longer than 9–12; wing broad with moderately short costa, costal ratio 0.44; subgenital plate anterior portion semi-circular with broad, ovoid lumen, posterior portion of main body as broad as anterior portion with 2 small sub-basal triangular points; spermatheca large (57.5 x 32.5 µm), ovoid, very heavily sclerotized, neck short, moderately stout, curved ( Figs. 26, 29–30 View FIGURES 22–30 )......................................... D. cyrtostyla View in CoL n. sp.
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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