Dissomphalus wusheanus Terayama, 2001

MUGRABI, DANIELE F. & AZEVEDO, CELSO O., 2013, Revision of Thai Dissomphalus Ashmead, 1893 (Hymenoptera, Bethylidae), with description of twenty four new species, Zootaxa 3662 (1), pp. 1-73 : 19-23

Dissomphalus wusheanus Terayama, 2001

(Figs (1–2, 22–23, 102, 151–154)

Dissomphalus wusheanus Terayama 2001: 87 (♂, description, figs 20-21); 2003: 13 (cladistic analysis).

Diagnosis. MALE. Mandible with three apical teeth. Clypeus with trapezoidal median lobe, median carina slightly almost straight and low in profile. Frons somewhat strongly coriaceous, frontal impression short, punctures shallow, dense and large. Vertex broad, slightly convex. Tergal process with pair of elliptical, very shallow lateral depression, latero-anterior border setosesetose, tubercle absent, tuft of few sparse short setae. Base of paramere wider than basiparamere apex; basiparamere narrow in dorsal view and wide in ventral view; aedeagal dorsal body compact, apex of inner lobe widely exposed, wider medially, lateral margin of aedeagal dorsal body slightly sinuous; ventral ramus narrower apically.

Description of metasoma. Metasomal tergite I with narrow and short groove. Metasomal tergite II with anterolateral margin not setosesetose; tergal process with pair of sub-lateral, transversely elliptical, shallow depressions, divergent posterad, outer lateral margin with short seta directed posterad ( Figs 1 , 22–23 ). Hypopygium with wide median stalk, apex truncate, 0.84 × as long as hypopygium, posterior margin almost straight ( Fig. 102 ). Genitalia ( Figs 151–154 ). Paramere with apex slightly curved medially, basal halves of dorsal and ventral margins parallel in lateral view, apical half inclined and straight in lateral view. Basivolsella with inner margin widened. Cuspis unusually wide. Digitus with expanded base. Aedeagus with ventral ramus much shorter than dorsal body, laminar, surface horizontal, dorsal surface concave, almost entirely wide, apex rounded; dorsal body with one pair of laminar outer apical lobes, surface vertical, apical half curved upward, apex medially scavated. Apodeme of aedeagus extending beyond genital ring.

FEMALE unknown.

Material examined. Holotype, ♂. TAIWAN. Wushe 1150 m, 23-III-1983, H. + M. Townes (PMAE). New material. THAILAND. Malaise trap, Chiang Mai: Doi Phahompok NP, Kewlom 1/montane forest, 2174 m, 1 ♂ ♂ T2815 , 1 ♂ T2821 ; Kiewlom 2: Montane Forest , 20°3.331’N 99°8.552’E, 2112 m, 3 ♂ 11-18.vii.2007, Komwuan Srisom leg., T2823 ; 7-14.ix.2007, Komwuan Srisom & Prasit Wongchai leg., 1 ♂ T2833 , 9 ♂ T2838 ; Doi Phaluang, 20°1.060’N 99°9.581’E, 1449 m, Wongchai. P. leg., 6 ♂ 20-27.vii.2007, T2928 , 1 ♂ 3-10.viii.2007, T2931 , 12 ♂ 13-20.vii.2007, T2846 ; 2 ♂ Headquarter 19°57.961’N 99°9.355’E, 569 m, 7-14.ii.2008, Seesom. K. leg., T2936 ; Doi Chiang Dao WS, 19°24.328’N 98°55.237’E, 31.vii-7.viii.2007, Songkrant Jagsu & Apichat Watwanich leg., 1 ♂ behind water reservior, 549 m, T2806 , 2 ♂ Nature Trail, 491 m, T2829 , 2 ♂ 28.viii-4.ix.2007, Thawatchai Boonthum & Anuchart Chaimuengchun leg., T2831 ; Huai Nam Dang NP, behind visitor house, 19°18.712’N 98°36.399’E, 1670 m, 6 ♂ 31.vii-7.viii.2007, T2804 , 24 ♂ 31.viii-7.ix.2007, T2811 ; Check Point 2, 19°18.239’N 98°36.292’E, 1590 m, 5 ♂ 31.vii-7.viii.2007, T2827 , 4 ♂ 31.viii-7.ix.2007, T2834 ; 2 ♂ Thung Buatong View Point, 19°17.470’N 98°36.033’E, 1464 m, 31.vii-7.viii.2007, T2849 ; Doi Inthanon NP, 2500 m, 18°35.361’N 98°29.157’E, Y. Areeluck leg., 1 ♂ summit forest, 15-22.vii.2006, T71 ; 1 ♂ summit marsh, 22.vii- 2.viii.2006, T118 ; 1 ♂ Kew Mae Pan, 18°33.163’N 98°28.800’E, 2200 m, 2-10.xi.2006, T388 ; campground pond, 18°32.657’N 98°31.482’E, 1200 m, 1 ♂ 8-15.vii.2006, T63 , 1 ♂ 2-9.viii.2006, T123 , 1 ♂ 6-13.ix.2006, T243 ; checkpoint 2, 18°31.559’N 98°29.941’E, 1700 m, 2 ♂ 29.vi-2.vii.2006, T44 , 4 ♂ 2-8.vii.2006, T56 , 1 ♂ 8- 15.vii.2006, T67 , 6 ♂ 22.vii-2.viii.2006, T121 , 2 ♂ 2-9.viii.2006, T127 , 1 ♂ 9-16.viii.2006, T180 , 3 ♂ 16- 24.viii.2006, T187 , 3 ♂ 30.viii-6.ix.2006, T238 , 1 ♂ 6-13.ix.2006, T244 , 6 ♂ 13-21.ix.2006, T250 , 3 ♂ 21- 27.ix.2006, T343 , 3 ♂ 27.ix-5.x.2006, T349 , 1 ♂ 5-12.x.2006, T365 , 1 ♂ 19-26.x.2006, T377 ; 18°31.554’N 98°29.940’E, 1700 m, 1 ♂ 24.xi-1.xii.2006, T1870 , 1 ♂ 1-8.xii.2006, T1876 , 1 ♂ 29.xii.2006 - 5.i.2007, T1897 , 1 ♂ 9-16.iii.2007, T1811 , 1 ♂ 15-22.iv.2007, T1845 ; 1 ♂ Campground Pond, 18°32.400’N 98°31.800’E, 1200 m, 15- 22.iv.2007, T1843 ; Vachiratharn Falls , 18°32.311’N 98°36.048’E, 700 m, 2 ♂ 29.vi-2.vii.2006, T40 , 1 ♂ 2- 8.vii.2006, T51 , 3 ♂ 8-15.vii.2006, T62 , 3 ♂ 15-22.vii.2006, T68 , 1 ♂ 22.vii-2.viii.2006, T116 , 2 ♂ 9-16.viii.2006, T175 , 1 ♂ 29.iv-6.v.2007, T1854 , 1 ♂ 8-15.v.2007, T1830 . Lampang: 2 ♂ Chae Son NP, Campground#3, 18°49.757’N 99°28.266’E, 487 m, 1-8.x.2007, Bunruen Kwunnui & Acharaporn Sukpeng leg., T2814 . Loei: Phu Ruea NP, 1 ♂ Nature trail, 17°30.740’N 101°20.650’E, 1353 m, 19-26.ix.2006, Nukoonchai Jaroenchai leg., T831 ; 2 ♂ Sa Sawan, 17°30.735’N 101°20.601’E, 1352 m, 5-12.iii.2007, Patikhom Tumtip leg., T2306 ; Nukoonchai Jaroenchai leg., 1 ♂ Pah Lo Noy, 17°30.502’N 101°20.868’E, 1343 m, 12-19.ix.2006, T827 ; 1 ♂ Huay Taey ditch, 17°30.128’N 101°20.339’E, 1233 m, 12-19.viii.2006, T529 ; Nern Wibaak ditch, 17°29.907’N 101°20.483’E, 1196 m, Patikhom Tumtip leg., 1 ♂ 19-26.viii.2006, T530 , 1 ♂ 5-12.ii.2007, T1705 , 1 ♂ 12-19.ii.2007, T1708 , 1 ♂ 26.ii-2.iii.2007, T1714 ; Nern Pitsawong, 17°29.676’N 101°21.093’E, 1168 m, 1 ♂ 12-19.xi.2006, T1120 , 1 ♂ 19- 26.xi.2006, T1123 , 1 ♂ 26.xi-2.xii.2006, T1126 ; 1 ♂ Ma Kraow ditch, 17°29.652’N 101°21.020’E, 1167 m, 19- 26.xi.2006, T1122 ; 2 ♂ Hua Dong Tham Sun, 17°29.540’N 101°20.995’E, 1130 m, 12-19.xi.2006, T1118 ; 1 ♂ Reservior, 17°28.826’N 101°21.330’E, 931 m, 19-26.i.2007, T1544 ; office, 17°28.805’N 101°21.242’E, 870 m, 1 ♂ 5-12.vii.2006, T312 , 12 ♂ 12-19.vii.2006, T315 ; 1 ♂ 26.vii-2.viii.2006, Nukoonchai Jaroenchai leg., T321 ; Subhnonghin, 17°28.772’N 101°21.308’E, 860 m, Patikhom Tamtip leg., 13 ♂ 5-12.vii.2006, T310 , 6 ♂ 12- 19.vii.2006, T313 ; 6 ♂ 19-26.vii.2006, Nukoonchai Jaroenchai leg., T316 ; Patikhom Tumtip leg., 4 ♂ Sab Nhong Hin, 17°28.772’N 101°21.308’E, 912 m, 5-12.i.2007, T1537 ; 1 ♂ Old road to the park, 17°27.901’N 101°21.301’E, 700 m, 5-12.xii.2006, T1263 ; 1 ♂ Behind checkpoint, 17°27.829’N 101°21.360’E, 691 m, 19- 26.xii.2006, T1268 ; Dry dipterocarp, 17°27.824’N 101°21.494’E, 668 m, 4 ♂ 12-19.xii.2006, T1264 , 8 ♂ 26.xii.2006 - 2.i.2007, T1270 ; 5 ♂ Hua Dong Tham Sun, 17°29.54’N 101°20.995’E 1130 m, 26.xi-2.xii.2006, Patikhom Tumtip leg. T1124 ; Phu Kradueng NP, 1 ♂ Hill evergren forest at Wang Gwang forest unit, 16°53.362’N 101°47.286’E, 1262 m, 28.xii.2006 - 3.i.2007, Sutin Gongla-sae leg., T1219 ; Savannah in pine forest, 16°53.092’N 101°47.413’E, 1257 m, Thanongsak Srisa-ad leg., 1 ♂ 9-16.i.2007, T1226 , 1 ♂ 16-23.i.2007, T1229 ; 1 ♂ Sutin Glong-lasae leg., Mixed deciduous forest behind the office, 16°56.589’N 101°42.074’E, 273 m, 14-21.x.2006, T782 ; Dry evergreen forest (Pong neep forest unit), 16°56.589’N 101°42.074’E, 273 m, 2 ♂ 21-28.x.2006, T786 , 1 ♂ 4-11.xi.2006, T792 ; 1 ♂ Sutin Khonglasae leg., Koke Hin Ngam, 16°51.958’N 101°50.668’E, 280 m, 9- 16.viii.2006, T482 ; 1 ♂ Huay Ta Hack, 16°51.958’N 101°50.668’E, 280 m, 16-23.viii.2006, T485 ; 3 ♂ Forest protection unit Loei.5 (Phakbung) 16°50.540’N 101°41.663’E, 406 m, 7-13.ii.2007, T1496 ; Noo Kerdlom leg., 2 ♂ 8-9.ii.2007, Pan trap, T1489 , Malaise trap, 10 ♂ 19-25.ii.2007, T1502 ; 4 ♂ 16°50.493’N 101°41.726’E, 412 m, 19-25.ii.2007, T1503 ; 2 ♂ 16°50.540’N 101°41.663’E, 406 m, Wuthicahi kwanjam leg., 13-19.ii.2007, T1499 ; 1 ♂ 16°50.463’N 101°41.687’E, 401 m, 13-19.ii.2007, T1498 ; 1 ♂ 25.ii-1.iii.2007, Sonkgran Kamtue leg., T1504 . Hom Waterfall, 17°7.411’N 104°1.016’E, 347 m, 1 ♂ 10-16.ix.2006, T611 , 1 ♂ 10-17.iii.2007, T2370 ; Winlon Khongnara leg., Kam Hom waterfall at Sao Hai area , 17°7.340’N 104°0.788’E, 344 m, 1 ♂ 4-10.ix.2006, T609 , 2 ♂ 10-16.ix.2006, T612 , 3 ♂ 17°7.415’N 104°1.179’E, 347 m, 16-22.ix.2006, T616 , 3 ♂ 17°7.340’N 104°0.788’E, 344 m, 16-22.ix.2006, T615 ; Behind forest protection unit at Huay Wien Prai, 1 ♂ 17°6.847’N 104°0.302’E, 376 m, 4-10.ii.2007, T1685 , 1 ♂ 17°6.810’N 104°0.318’E, 318 m, 10-17.ii.2007, T1687 ; Behind office, 17°3.543’N 103°58.452’E, 312 m, 1 ♂ 8-14.vii.2006, T197 , 1 ♂ 21-27.vii.2006, T203 ; 1 ♂ Sailom Tongboonchai leg., 15- 21.vii.2006, T200 ; 4 ♂ 17°3.521’N 103°58.450’E, 322 m, 21-27.vii.2006, T204 ; 1 ♂ 17°3.488’N 103°58.497’E, 318 m, 15-21.vii.2006, T202 ; Nam Hom Waterfall # Sao Hi, 17°7.340’N 104°0.788’E, 344 m, 2 ♂ 4-10.iii.2007, T2368 , 1 ♂ 17-25.iii.2007, T2374 ; North of well 17°3.543’N 103°58.452’E, 312 m, 6 ♂ 5-11.i.2007, T1516 , 8 ♂ 11-17.i.2007, T1519 , 1 ♂ 17-23.i.2007, T1522 , 8 ♂ 23-30.i.2007, T1525 ; West of well, 17°3.521’N 103°58.450’E, 322 m, 3 ♂ 5-11.i.2007, T1517 , 2 ♂ 11-17.i.2007, T1520 , 4 ♂ 17-23.i.2007, T1523 , 3 ♂ 23-30.i.2007, T1526 ; Behind national park office, 17°3.488’N 103°58.497’E, 318 m, 11-17.i.2007, 4 ♂ T1521 , 17-23.i.2007, 1 ♂ T1524 , 23-30.i.2007, 5 ♂ T1527 ; 1 ♂ Creek at northend of Huay Nam Pung Forest Unit, 16°55.789’N 103°54.158’E, 305 m, 13-19.xi.2006, Winlon Khongnara leg., T1099 ; Phu Pha Yon NP, Reservoir, 16°55.655’N 104°10.658’E, 280 m, Manop Ngoyjansri & Chatree Cheaukamjan, 1 ♂ 5-11.vii.2006, T290 , 1 ♂ 11-17.vii.2006, T293 ; Channel, 16°55.639’N 104°10.748’E, 295 m, 2 ♂ 5-11.vii.2006, T289 , 1 ♂ 11-17.vii.2006, T292 , 1 ♂ 17-23.vii.2006, T295 ; 1 ♂ Deciduous dipterocarp forest, 16°55.449’N 104°10.757’E, 295 m, T297 ; Phu Phan NP, 1 ♂ Forest near Huay Nam Pung Dam, 16°54.713’N 103°54.294’E, 289 m, 19-25.xi.2006, Sailom Tongboonchai leg., T1101 ; 1 ♂ Creek at entrance of Huay Nam Pung Forest unit, 16°54.630’N 103°54.266’E, 281 m, 13-19.xi.2006, Winlon Khongnara leg., T1097 . Phitsanulok: Thung Salaeng Luang NP, Mixed deciduous forest (Gang Sopa watrfall), 16°52.642’N 100°49.441’E, 481 m, Pan trap, Pongpitak Pranee leg., 1 ♂ 14-15.x.2006, T754 , 1 ♂ 16-17.x.2006, T756 ; Malaise trap, 2 ♂ Deciduous forest 16°50.699’N 100°51.266’E, 501 m, 8-15.iv.2007, Pongpitak & Pranee & Sathit leg., T2399 ; Moist evergreen, 16°50.641’N 100°52.894’E, 557 m, Pongpitak Pranee leg., 2 ♂ 11-18.viii.2006 T566 , 6 ♂ 18-25.viii.2006, T569 , 4 ♂ 25.viii-1.ix.2006, T572 , 1 ♂ 1-8.ix.2006, T575 ; Mixed deciduous forest, 16°50.563’N 100°51.757’E, 481 m, 1 ♂ 18-25.ix.2006, T930 , 1 ♂ 25.ix-2.x.2006, T933 , 2 ♂ 2-9.x.2006, T936 ; 16°50.319’N 100°51.792’E, 479 m, 7 ♂ 11-18.viii.2006, T568 , 1 ♂ 25.viii-1.ix.2006, T574 , 5 ♂ 1-8.ix.2006, T577 ; Dry evergreen forest, 16°50.277’N 100°52.917’E, 486 m, 1 ♂ 2-9.x.2006, T935 , Pongpitak & Pranee & Sathit leg., 1 ♂ 18-25.iii.2007, T2388 , 4 ♂ 8-15.iv.2007, T2397 ; 16°50.217’N 100°52.541’E, 580 m, Pongpitak Pranee leg., 4 ♂ 25.viii-1.ix.2006, T573 , 1-8.ix.2006, 1 ♂ T576 . Phetchabun: Nam Nao NP, Forest protection unit - Huay Pralard cave , Noopean Hongyothi leg., 1 ♂ 16°44.999’N 101°27.804’E, 715 m, 29.i-5.ii.2007, T1627 ; 16°44.986’N 101°27.874’E, 711 m, 15-22.i.2007, 1 ♂ T1623 , 3 ♂ T1622 ; 1 ♂ 29.i-5.ii.2007, Leng janteab leg., T1629 ; Tham Pra Laad Forest Unit, 16°44.986’N 101°27.874’E, 711 m, 7 ♂ 14-21.viii.2006, T428 ; 3 ♂ 21-28.viii.2006, Noopean Hongyothi leg., T431 ; Hell evergreen forest, 16°44.402’N 101°34.560’E, 883 m, 5 ♂ 20-27.xi.2006, Leng Jantiep leg., T1326 ; Noopien Hongyothee leg., 1 ♂ 11-18.xii.2006, T1335 , 1 ♂ 27.xi-4.xii.2006, T1328 , 4 ♂ 4-11.xii.2006, T1331 , 5 ♂ 11-18.xii.2006, T1334 , 16°44.371’N 101°34.549’E, 834 m, Leng Jantiep leg., 10 ♂ 20- 27.xi.2006, T1324 , 4 ♂ 27.xi-4.xii.2006, T1327 , 6 ♂ 4-11.xii.2006, T1330 ; 1 ♂ 11-18.xii.2006, Noopien Hongyothee leg., T1333 ; Checkpoint 16°43.695’N 101°33.797’E, 921 m, 2 ♂ Pan trap, 6-7.iii.2007, Noopean Hongyothi & Leng Janteab leg., T2282 ; Malaise trap, Leng Janteab leg., 4 ♂ 15-22.iii.2007, T2291 ; 4 ♂ Forest Checkpoint, 16°43.695’N 101°33.797’E, 921 m, 23-30.x.2006, T1011 ; Noopean Hongyothi leg., 16°43.693’N 101°33.836’E, 917 m, 5 ♂ 13-20.xi.2006, T1020 , 2 ♂ 30.x-6.xi.2006, T1015 , 1 ♂ 6-13.xi.2006, T1018 , 5 ♂ 8- 15.iii.2007, T2289 , 1 ♂ 15-22.iii.2007, T2292 , 3 ♂ 22-29.iii.2007, Leng Janteab leg., T2295 ; 16°43.687’N 101°33.754’E, 924 m, 6 ♂ 6-13.xi.2006, Noopean Hongyothi leg., T1016 , Leng Janteab leg., 3 ♂ 1-8.iii.2007, T2284 , 1 ♂ 15-22.iii.2007, T2290 , 10 ♂ 22-29.iii.2007, T2293 ; Helicopter landing ground, 3-10.vii.2006, Noopean Hongyothi leg., 4 ♂ 16°43.184’N 101°35.137’E, 875 m, T268 ; 16°43.156’N 101°35.118’E, 890 m, 1 ♂ T269 ; 1 ♂ Pan trap, 6-7.vii.2006, T264 ; Malaise trap, 1 ♂ 10-17.vii.2006, T272 , 1 ♂ 17-24.vii.2006, T275 ; 16°43.113’N 101°35.134’E, 889 m, 3 ♂ 10-17.vii.2006, T273 , 1 ♂ 17-24.vii.2006, T276 , 4 ♂ 24-30.vii.2006, T279 , 1 ♂ 1-8.i.2007, T1440 ; Khao Kho NP, Somchai Chatchumnan and Sa-ink Singtong leg., 1 ♂ Nursery, 16°52.581’N 101°8.060’E, 520 m, 26.x-2.xi.2006, T814 ; Mix deciduous, 3 ♂ 16°39.589’N 101°8.185’E, 168 m, 26.i-2.ii.2007, T1401 ; 16°39.572’N 101°8.194’E, 171 m, 1 ♂ 5-12.i.2007, T1391 , 1 ♂ 12-19.i.2007, T1394 , 2 ♂ 12-19.i.2007, T1393 ; Mix deciduous near office, 1 ♂ 16°39.587’N 101°8.134’E, 220 m, 26.ii-5.iii.2007, T1611 ; ♂ 16°39.550’N 101°8.123’E, 230 m, 12-19.vii.2006, T167 ; 1 ♂ 16°39.479’N 101°8.105’E, 220 m, 12-19.vii.2006, T166 ; Thung Salaeng Luang NP, Kaeng Wang Nam Yen, 16°37.531’N 100°53.745’E, 2 ♂ 16-22.xi.2006, T1159 , 1 ♂ 22-29.xi.2006, T1162 , 1 ♂ 29.xi-6.xii.2006, T1165 , 11 ♂ s 6-13.xii.2006, T1168 ; 16°37.178’N 100°53.504’E, 4 ♂ 16-22.xi.2006, T1158 , 1 ♂ 22-29.xi.2006, T1161 , 7 ♂ 29.xi-6.xii.2006, T1164 , 7 ♂ 6-13.xii.2006, T1167 ; 2 ♂ 16°36.587’N 100°53.395’E, 16-22.xi.2006, T1157 ; Gang Wang Nam Yen, 16°37.178’N 100°53.504’E, 706 m, 3 ♂ 31.v-7.vi.2007, T2087 , 3 ♂ 7-14.vi.2007, T2090 ; 1 ♂ 16°36.587’N 100°53.395’E, 753 m, 7-14.vi.2007, T2089 ; Pine forest Gang Wang Nam Yen, 16°36.284’N 100°53.128’E, 749 m, Pan trap, 1 ♂ 19-20.xii.2006, T1277 , 1 ♂ 18-19.vi.2007, T2054 ; Malaise trap ,, 5 ♂ 22-28.xii.2006 T1286 ; 16°35.789’N 100°52.286’E, 769 m, 3 ♂ 22- 28.xii.2006, T1285 ; Pongpitak and Sathit leg., 7 ♂ 4-11.i.2007, T1291 , 1 ♂ 22-29.vi.2007, T2062 , 1 ♂ 22- 28.xii.2006, T1287 ; 16°35.805’N 100°52.286’E, 726 m, 2 ♂ 28.xii.2006 - 4.i.2007, T1290 , 1 ♂ 4-11.i.2007, T1293 ; 16°34.569’N 100°53.155’E, 750 m, 22-29.i.2007, 1 ♂ T1583 , 16°34.360’N 100°52.827’E, 725 m, 16-22.i.2007, 1 ♂ T1579 , 29.i-4.ii.2007, 3 ♂ T1585 , 16°34.339’N 100°53.425’E, 758 m, 22-29.i.2007, 1 ♂ T1584 , 4-11.ii.2007, 3 ♂ T1590 ; Nam Nao NP, 3 ♂ Hell evergreen Forest, 16°44.402’N 101°34.560’E, 883 m, 27.xi-4.xii.2006, Noopien Hongyothee leg., T1329 . Sakon Nakhon: Phu Phan NP, Dry evergreen forest near house at 1567 station 16°48.628’N 103°53.591’E, 522 m, S. Kongnara leg., 3 ♂ 3-9.xii.2006, T1241 , 3 ♂ 9-15.xii.2006, T1244 , 2 ♂ 22- 29.xii.2006, T1250 ; Winlon Kongnara leg., 1 ♂ 10-16.vi.2007, T2496 , 1 ♂ 16-22.vi.2007, T2499 ; 16°48.627’N 103°53.511’E, 512 m, S. Kongnara leg., 2 ♂ 9-15.xii.2006, T1243 , 4 ♂ 22-29.xii.2006, T1249 ; Dry evergreen forest near office, 16°48.618’N 103°53.476’E, 526 m, 1 ♂ 3-9.xii.2006, T1242 , 2 ♂ 9-15.xii.2006, T1245 , 1 ♂ 22- 28.vi.2007, Winlon Kongnara leg., T2503 . Kamphaeng Phet: Mae Wong NP, Chong Yen, 16°5.968’N 99°6.472’E, 1306 m, Chumpol Piluk & Aram Inpuang leg., 12 ♂ 1-8.x.2007, T2816 ; 16 ♂ 1-3.xii.2007, T2822 ; 8 ♂ 3-10.ix.2007, T2812 . Chaiyaphum: Tat Tone NP, Dry dipterocarp forest near swamp at Sab somboon forest unit, 16°1.059’N 101°58.603’E, 674 m, Tawit Jaruphan leg., 1 ♂, 5-12.xi.2006, T1137 ; 1 ♂ 12-19.xi.2006, T1140 ; 1 ♂ 26.xi-3.xii.2006, T1146 ; 1 ♂ Lawn near Sab Somboon forest unit, 16°0.792’N 101°58.472’E, 648 m, 26.xi- 3.xii.2006, Tawit Jaruphan leg., T1145 ; 1 ♂ Chaiyapoom forest fire station, 16°0.738’N 102°1.342’E, 195 m, 12- 19.xii.2006, Tawit Jaruphan & Orawan Budsawong leg., T1371 ; 1 ♂ Dry Dipterocarp Forest , 15°59.037’N 102°2.103’E, 250 m, 21-28.vi.2006, Lumyai Ittichan leg., T36 ; Tawit Jaruphan & Orawan Budsawong leg, Phu hang sing, 15°58.723’N 102°2.231’E, 290 m, 1 ♂ 5-12.vii.2006, T220 ; 1 ♂ 12-19.vii.2006, T223 ;, Nursery near head water, 15°58.344’N 102°2.169’E, 257 m,., 1 ♂ 5-12.viii.2006, T547 ; 1 ♂ 26.ii-2.iii.2007, T1737 ; Lum pa tao head water/dry evergreen, 15°58.486’N 102°2.239’E, 270 m, 1 ♂ 12-19.ii.2007, T1730 ; 1 ♂ 26.ii-2.iii.2007, T1736 ; Tawit Jaruphan leg., 2 ♂ Next to Pa Eang waterfall, 15°57.657’N 101°54.724’E, 301 m, 26.x-3.xi.2006, T688 ; Entrance to Pa Eang waterfall, 15°57.520’N 101°54.442’E, 297 m, 3 ♂ 12-19.x.2006, T681 ; 1 ♂ 26.x- 3.xi.2006, T687 ; 4 ♂ Streamside at Tat Fah waterfall, 15°56.463’N 102°5.953’E, 242 m, 5-12.iii.2007 Tawit Jaruphan & Orawan Budsawong leg., T2346 ; Katae Sa-nog & Buakaw Adnafai leg., Pa Hin Ngam NP, %deciduous%, 1 ♂, 15°40.232’N 101°26.942’E, 398 m, 12-19.xii.2006, T1348 , 15°39.966’N 101°27.198’E, 357 m, 1 ♂ 19-23.xii.2006, T1352 ; 1 ♂ 26.xii.2006 - 2.i.2007, T1355 ; Thepana waterfall, Savana, 2 ♂ 15°38.948’N 101°25.625’E, 604 m, 19-25.x.2006, T666 ; Mixed deciduous Forest, 15°38.920’N 101°25.077’E, 614 m, 2 ♂ 1- 7.x.2006, T656 ; 3 ♂ 7-13.x.2006, T659 ; 3 ♂ 13-19.x.2006, T662 ; 1 ♂ 19-25.x.2006, T665 ; Dry evergreen Forest, 15°38.884’N 101°25.840’E, 605 m, 3 ♂ 1-7.x.2006, T658 ; 2 ♂ 7-13.x.2006, T661 ; 3 ♂ 13-19.x.2006, T664 ; 5 ♂ 19-25.x.2006, T667 ; car park at Tung Dok Grajeaw, 15°38.438’N 101°23.576’E, 780 m, 1 ♂ 6-12.vii.2006, T330 , 1 ♂ 12-18.vii.2006, T333 ; Thung Dok Kra Jeow in Dry evergreen, 15°38.438’N 101°23.576’E, 780 m, 14 ♂ 7- 13.i.2007, T1457 , 1 ♂ 19-25.i.2007, T1463 ; 15°38.391’N 101°23.609’E, 750 m, 4 ♂ creek at Tung Dok Grajeaw, 19-24.vii.2006, T335 ; Car park at Thung Dok Kra Jeow, 5 ♂ 1-7.i.2007, T1453 ; 1 ♂ 7-13.i.2007, T1456 ; 1 ♂ 13- 19.i.2007, T1459 ; Tung Dok Grajeaw, 15°38.208’N 101°23.556’E, 720 m, 3 ♂ 19-24.vii.2006, T337 ; 1 ♂ 24- 30.vii.2006, T340 ; Ecotone between mix deciduous/dry dipterocarp, 15°38.132’N 101°23.922’E, 698 m, 6 ♂ 5- 11.viii.2006, T441 ; 3 ♂ 11-18.viii.2006, T444 ; 700 m, 4 ♂ 5-11.viii.2006, T440 ; 4 ♂ 18-24.viii.2006, T446 ; 1 ♂ 24-30.viii.2006, T449 ; 1 ♂ 11-18.viii.2006, T445 ; 1 ♂ Dry dipterocarp, 15°38.099’N 101°23.921’E, 698 m, 24- 30.viii.2006, T451 ; Dry Dipterocarp forest at Lan Hin Nau, 15°37.683’N 101°23.323’E, 681 m, 2 ♂ 7-13.ix.2006, T850 ; 1 ♂ 13-19.ix.2006, T853 ; 3 ♂ 19-26.ix.2006, T856 ; 15°37.540’N 101°23.313’E, 673 m, 3 ♂ 1-7.ix.2006, T846 ; 1 ♂ 7-13.ix.2006, T849 ; 1 ♂ 13-19.ix.2006, T852 ; 1 ♂ 19-26.ix.2006, T855 . Ubon Ratchathani: Pha Taem NP, Thongcome & Pakdee leg., 2 ♂ Don Huay Can 15°40.016’N 105°30.502’E, 246 m, 25.xii.2006 - 1.i.2007, 15°39.989’N 105°30.468’E, 238 m, 1-7.i.2007, T1475 ; 1 ♂ Huay Pok waterfall, 15°37.321’N 105°36.982’E, 419 m, 18-25.iv.2007, Bunlu Sapsiri leg., T2171 ; Irrigation area west of Huay Pok forest unit, 15°37.321’N 105°36.982’E, 419 m, 6-13.x.2006, 2 ♂ T719 ; 1 ♂ T720 ; 1 ♂ 13-20.x.2006, T722 ; 1 ♂ 20-27.x.2006, T726 ; 1 ♂ 27.x-3.xi.2006, T728 ; Pornthip Tonsu leg., Huay Pok waterfall, 1 ♂ T721 ; 3 ♂ 27.x-3.xi.2006, T730 ; 1 ♂ west of HuayPok substation, 15°37.212’N 105°36.903’E, 438 m, 11-18.iv.2007, Bunlu Sapsiri leg., T2167 ; Kee thao cave, 15°32.001’N 105°35.857’E, 157 m, Mitchai Sai-ngam leg., 1 ♂ 4-11.ix.2006, T887 ; 1 ♂ 25.ix-1.x.2006, T896 ; Huay Sa Nhom plateau, 15°27.435’N 105°34.838’E, 238 m, Sorawit and Thongdee leg., 1 ♂ 4-11.xi.2006. T1052 , 3 ♂ 11-18.xi.2006, T1055 ; 1 ♂ First level-Huay Sa-nhom waterfall, 15°27.407’N 105°34.867’E, 230 m, 9- 16.v.2007, Sorawit Mingman leg., T2188 ; Wild flower field 1, 15°27.336’N 105°34.870’E, 232 m, Sorawit and Thongdee leg., 4 ♂ 18-25.xi.2006, T1060 ; 2 ♂ 25.xi-2.xii.2006, T1063 . Suphanburi: 5 ♂ Pu Toei NP, Huai Mongpae/red road, 14°56.985’N 99°26.780’E, 300 m, 16-23.vii.2008, Saunbua.L. leg., T3117 . Kanchanaburi: Khuean Srinagarindra NP, 1 ♂, Tha Thung-na/Chong Kraborg, 14°38.123’N 98°59.657’E, 210 m, 9-16.x.2008, Boonnam & Phumarin leg., T3469 ; Huay Mae Kamint, Somboon & Daorueng leg., 1 ♂ Tourist center, 14°38.312’N 98°59.643’E, 210 m, 14-21.v.2009, T4746 ; 50 m / SW of Tourist center, 14°29.972’N 98°53.035’E, 8 ♂ 18-25.ix.2008, T3443 , 2 ♂ 25.ix-2.x.2008, Chatchawan & Boonkam leg., T3444 ; 3 ♂ Behind tourist center, 14°38.155’N 98°59.85’E, 210 m, 15-22.viii.2008, Chatchawan leg., T3420 ; 1 ♂ Tourist center, 14°38.136’N 98°59.837’E, 210 m, 11-18.vii.2008, Somboon & Chatchawan leg., T3432 ; 1 ♂ Tha Thung-na/Chong Kraborg, 14°29.972’N 98°53.035’E, 210 m, 25.xii.2008 - 1.i.2009, Boonnam & Phumarin leg., T4437 . Nakhon Ratchasima: Khao Yai NP, 3 ♂, Dong Ngu How area , 14°28.524’N 101°22.928’E, 757 m, 26.xii.2006 - 2.i.2007, Wirat Sook kho leg., T1312 ; Pong Sandow leg., 2 ♂ Moist evergreen forest at Dong Suer Paan, 14°27.511’N 101°22.408’E, 760 m, 5-12.xii.2006, T1305 ; 1 ♂ evergreen forest near Tiger Trail, 14°27.511’N 101°22.408’E, 760 m, 26.vi-2.vii.2007, T2232 . Nakhon Nayok: Khao Yai NP, Wirat Sookkho leg., Moist evergreen forest near the reservoir, 14°27.119’N 101°21.482’E, 699 m, 1 ♂ 5-12.xi.2006, T991 ; 5 ♂ 12-19.xi.2006, T994 ; 1 ♂ 19-26.xi.2006, T997 ; 1 ♂ 26.xi- 3.xii.2006, T1000 ; 1 ♂ behind Training Center, 14°27.119’ N 101°21.482’E, 699 m, 12-19.v.2007, T2268 ; entrance of Nong Pak Chee Trail, 14°27.115’N 101°21.951’E, 733 m, 1 ♂ 5-12.v.2007, T2264 ; 1 ♂ 12-19.v.2007, Pong Sandao leg., T2267 ; 1 ♂ 26.v-2.vi.2007, T2273 ; Pong Sandao leg., Lam Takong view point, 14°25.820’N 101°23.754’E, 744 m, 1 ♂ 19-26.x.2006, T748 ; 1 ♂ 26.x-2.xi.2006, T751 ; 1 ♂ 14°25.565’N 101°23.442’E, 726 m, 26.iv-2.v.2007, T2130 ; 14°24.515’N 101°22.432’E, 750 m, 1 ♂ Nature trail insecondary moist evergreen forest, 5.vii-12.viii.2006, T400 ; 2 ♂ near Training Center 2, 5-12.ii.2007, T2243 ; Khao Keow spirit house, 14°22.960’N 101°23.253’E, 750 m, 1 ♂ 12-19.ix.2006, Wirat Sook-kho leg., T912 ; 1 ♂ 26.ix-3.x.2006, Pong Sandao leg., T916 . Petchaburi: Kaeng Krachan NP, Panernthung, 1 ♂ km30, 12°49.598’N 99°21.827’E, 19-26.ix.2008, Sirichai & Jatupon leg., T4369 ; 2 ♂ km27, 12°49.302’N 99°22.263’E, 4-11.xii.2008, Sirichai leg., T4399 ; km33/ helipad, 12°50.177’N 99°20.688’E, 735 m, Sirichai leg., 1 ♂ 4-11.xii.2008, T4393 , 2 ♂ 11-18.ii.2009, T4415 ; 1 ♂ Pa La-U/ Huai Palao Forest Unit 3, 12°32.149’N 99°28.265’E, 28.xii.2008 - 4.i.2009, Thongbai leg., T4558 .

Variations. Setae of metasomal tergite II short or long, thick or thin, placed lateromedially; depressions of tergal process fully rounded or rounded with truncate apical margin or sub-circular, shallow to deep, touching posterior margin of tergite I, touching or not touching posterior margin of tergite II with tufts of depression with elongate to circular; seta length very short to long; outer lateral margin of depression with long and thin seta; inner lateral surface of depression with small or wide tubercle, inclined laterad, with small and narrow pit.

Remarks. Dissomphalus wusheanus was the most abundant species in the collected material, representing 42% of the specimens. D. wusheanus was previously known only in Taiwan, where only three specimens had been recorded ( Terayama 2001). As a result of the present study, it has also been identified in Thailand, which might point to a potentially wider scope of distribution encompassing the neighboring countries. Upon visiting South China Agricultural University, the second author found hundreds of specimens of this species collected from Hainan, China, which strengthens the hypothesis postulating a wider area of distribution. Dissomphalus wusheanus is similar to D. ferrurus sp. nov. because of the following characteristics: compact appearance of aedeagal dorsal body with inner lobe widely exposed, wider medially; and basiparamere narrow in dorsal view and wide in ventral view. However, in D. wusheanus , the base of paramere is wider than basiparamere apex; the basivolsella has no projection; the lateral margin of aedeagal dorsal body is slightly sinuous; and the ventral ramus is narrow apically.

Distribution. Thailand (Chiang Mai, Lampang, Sakon Nakhon, Phitsanulok, Phetchabun, Sakon Nakhon,

Figures

FIGURES 1–8. 1–2, Dissomphalus wusheanus, 1, metasoma in dorsal view, 1a, metasomal tergite II length; 1b distance between the metasomal tergite II depressions; 2, hypopygium in dorsal view, 2a hypopygium length; 2b, hypopygium median stalk length; 2c hypopygium lateral stalk length; 3–7 metasoma in dorsal view, 3, D. brevinervis; 4, D. chiangmaiensis; 5, D.

FIGURES 9–23. Metasoma, 9–11, Dissomphalus browni, 9, metasoma in dorsal view; 10, tergal process of metasoma in dorsal view; 11, tubercle of tergal process in dorsal view; 12–18, D. chiangmaiensis, 12, metasoma in dorsal view; 13, tergal process of metasoma in dorsal view; 14, tubercle of tergal process in dorsal view; 15, tubercle of tergal process in lateral view; 16, metasoma in dorsal view; 17, tergal process of metasoma in dorsal view; 18, pit of tregal process in dorsal view; 19–21, D.

FIGURES 95–106. Hypopygium in dorsal view, 95, Dissomphalus. brevinervis; 96, D. browni; 97, D. chiangmaiensis; 98, D.

FIGURES 148–158. Male genitalia, 148–150, Dissomphalus thaianus, 148, dorsal view; 149, apex of aedeagus in dorsal view; 150, ventral view; 151–154, D. wusheanus; 151, dorsal view; 152, lobe of dorsal body of aedeagus in dorsal view; 153, ventral