Dissomphalus verus, MUGRABI, DANIELE F. & AZEVEDO, CELSO O., 2013
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publication ID |
https://doi.org/ 10.11646/zootaxa.3662.1.1 |
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publication LSID |
lsid:zoobank.org:pub:F8E5DC33-8045-4511-B046-90F59E97BD1D |
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persistent identifier |
https://treatment.plazi.org/id/E004C965-FD3E-6018-FF1A-FF00FA535800 |
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treatment provided by |
Felipe |
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scientific name |
Dissomphalus verus |
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Key for Dissomphalus males from the Oriental region
1. Depression of tergal process with outer lateral margin higher than inner lateral margin ( Fig. 25 View FIGURES 24–38 ); aedeagal dorsal body with two pairs of outer apical lobes ( Fig. 166 View FIGURES 159–167 )....................................................................... 2
- Depression of tergal process with outer and inner lateral margins equally high ( Fig. 13 View FIGURES 9–23 ); aedeagal dorsal body with one pair of outer apical lobes ( Fig. 133 View FIGURES 128–136 )............................................................................. 8
2(1). Posterior margin of hypopygium straight with three small concavities ( Fig. 103 View FIGURES 95–106 ).................................... 3
- Posterior margin of hypopygium concave or with median concavity ( Fig. 104 View FIGURES 95–106 )..................................... 4
3(2). Outer lateral margin of dorsal outer lobe of aedeagal dorsal body with laminar process ( Fig. 160 View FIGURES 159–167 )......... D. verus sp. nov.
- Outer lateral margin of dorsal outer lobe of aedeagal dorsal body without laminar process ( Fig. 155 View FIGURES 148–158 )....... D. epitus sp. nov.
4(2). Posterior margin of hypopygium with angulate corners ( Fig. 106 View FIGURES 95–106 )............................................... 5
- Posterior margin of hypopygium with rounded corners ( Fig. 95 View FIGURES 95–106 )................................................. 6
5(4). Dorsal outer lobe of aedeagal dorsal body with sickle-shaped laminar process ( Fig. 163 View FIGURES 159–167 ), with apical margin strongly concave, turned laterad; apex of aedeagal ventral ramus as high as apex of volsella ( Fig. 164 View FIGURES 159–167 ).................... D. prilus sp. nov.
- Dorsal outer lobe of aedeagal dorsal body with transversely elliptical laminar process, turned mesad; apex of aedeagal ventral ramus as high as apex of basivolsella ( Fig. 167 View FIGURES 159–167 )................................................ D. daneus sp. nov.
6(4). Ventral outer lobe of aedeagal dorsal body strongly concave ( Fig. 131 View FIGURES 128–136 ), apex laterad................. D. browni Terayama
- Ventral outer lobe of aedeagal dorsal body straight or nearly so ( Fig. 143 View FIGURES 137–147 ), apex dorsad.............................. 7
7(6). Aedeagal ventral ramus very wide ( Fig. 147 View FIGURES 137–147 ); aedeagal apodeme strongly dilated ( Fig. 145 View FIGURES 137–147 ); ratio between aedeagal ventral ramus and dorsal body 1:3.5........................................................ D. philippinensis Terayama
- Aedeagal ventral ramus narrow ( Fig. 144 View FIGURES 137–147 ); aedeagal apodeme non-dilated ( Fig. 141 View FIGURES 137–147 ); ratio between aedeagal ventral ramus and dorsal body 1:2.5.................................................................... D. nepalensis Terayama
8(1). Basivolsella with projection ( Fig. 169 View FIGURES 168–176 )..................................................................... 9
- Basivolsella without projection ( Fig. 187 View FIGURES 186–193 )................................................................. 16
9(8). Aedeagal dorsal body evenly wide ( Fig. 171 View FIGURES 168–176 ) or base slightly wider than remainder of dorsal body ( Fig. 183 View FIGURES 177–185 )............ 10
- Aedeagal dorsal body wider medially ( Fig. 168 View FIGURES 168–176 )............................................................ 11
10(9). Paramere with ventral surface deeply divided ( Fig. 174 View FIGURES 168–176 ); aedeagal dorsal body evenly wide ( Fig. 171 View FIGURES 168–176 ); base of basivolsella with two short and basally articulated projections ( Fig. 173 View FIGURES 168–176 ); aedeagal ventral ramus 0.5x as long as dorsal body................................................................................................. D. gionus sp. nov.
- Paramere with ventral surface undivided ( Fig. 185 View FIGURES 177–185 ); base of aedeagal dorsal body slightly wider than remainder of dorsal body; median region of basivolsella with two long projections, medially articulated ( Fig. 184 View FIGURES 177–185 ); aedeagal ventral ramus almost as high as dorsal body........................................................................... D. davus sp. nov.
11(9). Median stalk of hypopygium shorter than hypopygium width, its apex truncate (Fig. 107); apex of paramere angulate or subangulate in dorsal view ( Fig. 168 View FIGURES 168–176 ); basivolsella with one projection ( Fig. 169 View FIGURES 168–176 )..................................... 12
- Median stalk of hypopygium as long as or longer than hypopygium width, its apex rounded (Fig. 114); apex of paramere rounded in dorsal view ( Fig. 175 View FIGURES 168–176 ); basivolsella with more than one projection ( Fig. 176 View FIGURES 168–176 )............................ 14
12(11). Median part of genital ring with inclination angle of about 45º on dorsal view; paramere with apical constriction on dorsal view( Fig. 168 View FIGURES 168–176 ); basivolsella with medially articulated projection; cuspis with one ramus ( Fig. 169 View FIGURES 168–176 )....... D. gabrus sp. nov.
- Median part of genital ring with an inclination angle between 45 and 90º in dorsal view; paramere without apical constriction in dorsal view ( Fig. 179 View FIGURES 177–185 ); basivolsella with basally articulated projection; cuspis with two rami ( Fig. 181 View FIGURES 177–185 ).............. 13
13(12). Corner of posterior margin of hypopygium angulate, with short median evagination (Fig. 110); median region of inner lateral margin of basiparamere with long projection; basivolsella with apical projection ( Fig. 181 View FIGURES 177–185 ); aedeagal ventral ramus almost as long as dorsal body....................................................................... D. julius sp. nov.
- Corner of posterior margin of hypopygium rounded, not medially evaginated ( Fig. 127 View FIGURES 119–127 ); median region of inner lateral margin of basiparamere without projection; basivolsella with median projection ( Fig. 222 View FIGURES 218–222 ); aedeagal ventral ramus much shorter than dorsal body........................................................................... D. ferrurus sp. nov.
14(11). Lateral margins of hypopygium slightly convergent posterad; apex of hypopygeal median stalk without constriction (Fig. 109); apex of aedeagal dorsal body not surpassing paramere apex ( Fig. 175 View FIGURES 168–176 ); ventral ramus with non-sinuous surface ( Fig. 176 View FIGURES 168–176 ).. 15
- Lateral margins of hypopygium strongly convergent posteriorly; apex of hypopygeal median stalk with small constriction (Fig. 114); apex of aedeagal dorsal body aligned with paramere apex ( Fig. 192 View FIGURES 186–193 ); ventral ramus with sinuous surface ( Fig. 193 View FIGURES 186–193 )........................................................................................... D. joelus sp. nov.
15(14). Paramere with ventral surface deeply divided ( Fig. 177 View FIGURES 177–185 ), apical margin obliquely straight ( Fig. 175 View FIGURES 168–176 ); basivolsella with three projections ( Fig. 176 View FIGURES 168–176 ); aedeagal dorsal body wide, its lateral margins strongly convex ( Fig. 175 View FIGURES 168–176 )........ D. cherrus sp. nov.
- Paramere with ventral surface undivided, apical margin slightly convex in ventral view; basivolsella with two projections ( Fig. 203 View FIGURES 202–209 ); aedeagal dorsal body narrow, its lateral margins straight ( Fig. 202 View FIGURES 202–209 )............................ D. turinus sp. nov.
16(15). Lateral surface of aedeagal dorsal body with warts ( Fig. 133 View FIGURES 128–136 ).................................................. 17
- Lateral surface of aedeagal dorsal body without warts ( Fig. 148 View FIGURES 148–158 )............................................... 18
17(16). Hypopygeal median stalk of hypopygium with straight margin, apex truncate ( Fig. 97 View FIGURES 95–106 ); apex of inner lateral margin of aedeagal ventral ramus concave ( Fig. 136 View FIGURES 128–136 )................................................ D. chiangmaiensis Terayama
- Hypopygeal median stalk of hypopygium with convex margin, apex angulate ( Fig. 98 View FIGURES 95–106 ); apex of inner lateral margin of aedea- 19(18). Aedeagal apodeme extremely long, strongly converging basad................................. D. thaianus Terayama
- Aedeagal apodeme not as long as above, parallel or slightly converging basad..................................... 20
20(19). Base of aedeagal ventral ramus about as wide as medially ( Fig. 187 View FIGURES 186–193 )............................................ 21
- Aedeagal ventral ramus much narrower basally than medially ( Fig. 189 View FIGURES 186–193 )......................................... 23
21(20). Aedeagal dorsal body progressively narrowing apicad ( Fig. 186 View FIGURES 186–193 ); hypopygeal median stalk wide (Fig. 112); basiparamere evenly narrow on dorsal view; paramere with base wider than basiparamere apex, undivided ventral surface............. 22
- Aedeagal dorsal body narrowing sharply apicad ( Fig. 212 View FIGURES 210–217 ); hypopygeal median stalk narrow ( Fig. 122 View FIGURES 119–127 ); basiparamere irregularly narrow on dorsal view; paramere with base as wide as basiparamere apex, divided ventral surface.... D. geanus sp. nov.
22(21). Median region of genital ring with inclination angle between 45 and 90º in dorsal view; aedeagus dorsal body wide, lateral margin strongly convex medially ( Fig. 186 View FIGURES 186–193 ); aedeagal ventral ramus evenly wide ( Fig. 187 View FIGURES 186–193 )............. D. zethus sp. nov.
- Median region of genital ring with inclination angle of about 45º in dorsal view; aedeagal dorsal body narrow, lateral margin slightly convex ( Fig. 194 View FIGURES 194–201 ); aedeagal ventral ramus wider medially ( Fig. 195 View FIGURES 194–201 )........................ D. nandus sp. nov.
23(20). Aedeagal ventral ramus shorter than dorsal body............................................................ 24
- Aedeagal ventral ramus as long as dorsal body............................................................. 25
24(23). Dorsal surface of basiparamere almost as wide as ventral surface; aedeagal dorsal body narrow ( Fig. 204 View FIGURES 202–209 ); aedeagal ventral ramus much wider apically, with three pairs of long projections ( Fig. 206 View FIGURES 202–209 )........................... D. paulus sp. nov.
- Dorsal surface of basiparamere much narrow than ventral surface; aedeagal dorsal body wide ( Fig. 208 View FIGURES 202–209 ); aedeagal ventral ramus wider medially, without projections ( Fig. 209 View FIGURES 202–209 )........................................... D. barbus sp. nov.
25(23). Ventral surface of paramere slightly divided ( Fig. 190 View FIGURES 186–193 ); apex of aedeagal ventral ramus angulate.......... D. hetus sp. nov.
- Ventral surface of paramere undivided ( Fig. 185 View FIGURES 177–185 ); apex of aedeagal ventral ramus rounded.......................... 26
26(25). Aedeagal ventral ramus constricted medially ( Fig. 220 View FIGURES 218–222 ); aedeagal dorsal body entirely wide ( Fig. 218 View FIGURES 218–222 )... D. mugrus sp. nov.
- Aedeagal ventral ramus wide medially ( Fig. 129 View FIGURES 128–136 ); aedeagal dorsal body irregularly wide ( Fig. 128 View FIGURES 128–136 ).................... 27
27(26). Clypeus with one or three median teeth; hypopygeal median stalk ( Fig. 102 View FIGURES 95–106 ); aedeagal dorsal body slightly narrow subapically ( Fig. 151 View FIGURES 148–158 )........................................................................................... 28
- Clypeus without median teeth; hypopygeal median stalk narrow ( Fig. 95 View FIGURES 95–106 ); aedeagal dorsal body with abruptly narrow subapically ( Fig. 128 View FIGURES 128–136 )...................................................................... D. brevinervis Kieffer
28(27). Hypopygeal posterior margin concave ( Fig. 123 View FIGURES 119–127 ); aedeagal apodeme with apex turned mesad ( Fig. 214 View FIGURES 210–217 ); aedeagal ventral ramus longer than wide ( Fig. 215 View FIGURES 210–217 )........................................................ D. magnetus sp. nov.
- Hypopygeal posterior margin straight ( Fig. 102 View FIGURES 95–106 ); aedeagal apodeme with apex turned mesad ( Fig. 151 View FIGURES 148–158 ); aedeagal ventral ramus almost as long as wide ( Fig. 153 View FIGURES 148–158 )...................................................... D. wusheanus Terayama
29(18). Hypopygeal median stalk wide (Fig. 116); basiparamere evenly narrow ( Fig. 196 View FIGURES 194–201 ); cuspis with two rami ( Fig. 197 View FIGURES 194–201 )....... 30
- Hypopygeal median stalk narrow ( Fig. 121 View FIGURES 119–127 ); basiparamere irregularly wide ( Fig. 210 View FIGURES 210–217 ); cuspis with one ramus ( Fig. 211 View FIGURES 210–217 ) … 31
30(29). Hypopygium sub-trapezoidal, apex of median stalk not bifurcated (Fig. 116); inner lobes of aedeagal dorsal body widely visible in ventral view and not visible in dorsal view ( Fig. 197 View FIGURES 194–201 )....................................... D. jubus sp. nov.
- Hypopygium rounded, apex of median stalk bifurcated (Fig. 117); inner lobes of aedeagal dorsal body not visible in dorsal or ventral views............................................................................ D. kelsus sp. nov.
31(29). Hypopygeal of posterior margin without median evagination ( Fig. 121 View FIGURES 119–127 ); digitus extremely long, almost as long as paramere; aedeagal ventral ramus wider than long ( Fig. 211 View FIGURES 210–217 )............................................... D. robus sp. nov.
- Hypopygeal posterior margin with median evagination projected toward dorsal region ( Figs 124–125 View FIGURES 119–127 ); digitus regular-sized; aedeagal ventral ramus longer than wide ( Fig. 217 View FIGURES 210–217 )............................................. D. lidinus sp. nov.
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