Leptotrombidium akamushi ( Brumpt, 1910 )

Stekolnikov, Alexandr A., 2021, A checklist of chigger mites (Acariformes: Trombiculidae) of Southeast Asia, Zootaxa 4913 (1), pp. 1-163 : 111

Leptotrombidium akamushi ( Brumpt, 1910)

Trombidium akamushi Brumpt, 1910: 506 , fig. 335.

Trombicula akamushi: Gater, 1932: 146 ; Womersley & Heaslip 1943: 84, pl. III (5); Fuller 1952: 23; Gunther 1952: 10.

Trombicula (Trombicula) akamushi: Thor & Willmann 1947: 265 , fig. 324 (larva), 346 (nymph).

Trombicula (Leptotrombidium) akamushi: Womersley 1952: 67 ; Wharton & Fuller 1952: 51; Sasa & Jameson 1954: 292, fig. 27; Traub & Audy 1954a: 74, figs. 79–86; Audy 1957: 227; Womersley & Audy 1957: 254; Chen & Hsu 1958a: 411.

Leptotrombidium akamushi: Radford 1954: 260 ; Stekolnikov 2013: 65 .

Leptotrombidium (Leptotrombidium) akamushi: Lakshana 1973: 6 ; Vercammen-Grandjean 1969: 98, figs. on p. 99; Vercammen-Grandjean & Langston 1976: 261, pls. 17, 18; Wen 1984b: 304; Li et al. 1997: 107, fig. 2-1-2; Chau et al. 2007: 35, figs. 12, 13; Chung et al. 2015a: 1249; 2015b: 45, fig. 11.

Microtrombidium brumpti Hirst, 1915b: 82 .

Type deposition. L. akamushi: UCDC (neotype); M. brumpti : no data.

Type data. Ex Microtus montebelli , Japan, Akita Prefecture .

Hosts. RODENTIA : Apodemus agrarius , A. semotus , Atherurus macrourus , Bandicota indica , Cavia porcellus , Leopoldamys edwardsi , Menetes berdmorei , Microtus montebelli , Mus caroli , M. musculus , Niviventer andersoni , N. coninga , Rattus andamanensis , R. argentiventer , R. exulans , R. losea , R. mordax , R. norvegicus , R. rattus , R. tanezumi , R. tiomanicus ; SCANDENTIA : Tupaia glis , T. minor , T. montana ; SORICOMORPHA : Crocidura dsinezumi , C. tanakae , Suncus murinus ; LAGOMORPHA : Leporidae gen. sp.; CARNIVORA : Canis lupus familiaris , Felis catus , Herpestes javanicus exilis ; ARTIODACTYLA : Bos taurus , Bubalus bubalis , Tragulus javanicus ; PRIMATES: Homo sapiens, Simiiformes gen. sp., PERAMELOMORPHIA: Echymipera kalubu , E. kalubu cockerelli ; SQUAMATA : Eutropis multifasciata ; AVES: Acrocephalus orientalis , Alauda arvensis japonica , Asio flammeus , Calidris ruficollis , Centropus bengalensis javanensis , C. sinensis bubutus , Excalfactoria chinensis chinensis , Francolinus pintadeanus , Gallirallus striatus gularis , Gallus gallus domesticus , Lanius bucephalus , Phasianus versicolor , Porzana cinerea , Streptopelia orientalis , Turnix suscitator atrogularis , T. suscitator taigoor .

Distribution. Burma, Ceylon, China, Japan, Korea, Malaysia, Papua New Guinea, Philippines, Taiwan, Thailand, Vietnam.

Remarks. 1. Trombicula fletcheri Womersley and Heaslip, 1943 and T. obscura Womersley, 1944 were synonymized with T. akamushi by Womersley (1952), but these acts were not generally accepted ( Vercammen-Grandjean & Langston 1976).

2. I have not listed here old unavailable names, namely, Kedania tanakai Kishida, 1917 and Trombicula coarctata Kitashima and Miyajima, 1918 , which were included in the lists of synonyms for L. akamushi previously ( Gunther 1952; Fuller 1952; Wharton & Fuller 1952).

3. Vercammen-Grandjean (1969) established that any type material of Trombidium akamushi does not exist and designated a neotype, which was termed by him as “ lectotype ” ( Vercammen-Grandjean 1969; Vercammen-Grandjean & Langston 1976).

4. Vercammen-Grandjean & Langston (1976) questioned the presence of L. akamushi in Southeast Asia and noted that its range does not overlap with the range of L. deliense , according to critically revised data. Consequently, its list of hosts should in reality be much lesser than given above.