PIMOIDAE WUNDERLICH, 1986

Hormiga, Gustavo, Buckle, Donald J. & Scharff, Nikolaj, 2005, Nanoa, an enigmatic new genus of pimoid spiders from western North America (Pimoidae, Araneae), Zoological Journal of the Linnean Society 145 (2), pp. 249-262 : 252

PIMOIDAE WUNDERLICH, 1986

Pimoinae. – Wunderlich (1986: 16)

Pimoidae . – Hormiga (1993), type genus by monotypy Pimoa Chamberlin & Ivie. See also Hormiga (1994a, 2003).

Diagnosis: Male pimoids are distinguished form other araneoid spiders by the following combination of characters: palpus with integral paracymbium (intersegmental in Weintrauboa ), a retrolateral cymbial sclerite, a dorsoectal cymbial process, and cuspules (modified macrosetae) on either the cymbial process ( Pimoa ; Hormiga, 1994a: figs 11 and 68; Nanoa , Fig. 1A–B, D ) or the dorsal surface of the cymbium ( Weintrauboa ; Hormiga, 2003: figs 1e, 5e). Conductor and median apophysis present in most species. Embolus continuous with the tegulum (the typical linyphiid embolic division is absent), with an embolic process of varying morphology (absent in Nanoa ). The epigynum is protruding (except in Nanoa ), with a dorsal to lateral fold or groove with the copulatory opening at the distal end ( Hormiga, 1994a: figs 14, 414; Figs 3C , 5A ); fertilization ducts are anteriorly ( Pimoa ), posteriorly ( Nanoa ) or mesally ( Weintrauboa ) orientated. As in linyphiids, pimoids have stridulatory striae on the ectal side of the chelicerae (but the striae are absent in Weintrauboa and Nanoa ), build sheet-webs and exhibit autospasy at the patellatibia junction.

Description: See Hormiga (1994a) for family description.

Phylogenetics: The monophyly of Pimoidae is supported by the following four unambiguous putative synapomorphies: a dorsoectal cymbial process; cymbial cuspules (modified macrosetae); a retrolateral cymbial sclerite (pimoid cymbial sclerite, PCS); and the embolic process (pimoid embolic process, PEP, lost in Nanoa ).

Composition: Three genera, Pimoa Chamberlin & Ivie , Weintrauboa Hormiga and Nanoa gen. nov.

Figures

Figure 1. Nanoa enana sp. nov., male palp (embolus in a slightly displaced position; normally its distal end rests tightly against the tegulum, next to the conductor). A, ventral; B, dorsoectal (arrow points out to homologue of alveolar sclerite); C, mesal; D, ectal. Scale bars = 0.1 mm. See main text for abbreviations.

Figure 3. Nanoa enana sp. nov. A, male, ventral; B, female, dorsal; C, epigynum, ventral; D, cleared epigynum, ventral (arrow points out to possible location of copulatory opening); E, cleared epigynum, dorsal (arrow points out to distal end of fertilization duct). Scale bars: A, B = 0.5 mm; C–E = 0.1 mm. See main text for abbreviations.

Figure 5. Nanoa enana sp. nov. Female. A, epigynum, ventral; B, epigynum, caudoventral; C, epigynum, caudal; D, partially damaged PLS and PMS (PMS cylindrical spigot has shaft broken off; left PLS, on right, has one cylindrical spigot and part of the triplet broken off); E, right PLS triplet with ectal cylindrical spigot; F, prosoma, lateral. See main text for abbreviations.