Neoperla leopoldina Navás, 1932

Zwick, Peter & Zwick, Andreas, 2023, Revision of the African Neoperla Needham, 1905 (Plecoptera: Perlidae: Perlinae) based on morphological and molecular data, Zootaxa 5316 (1), pp. 1-194 : 185-191

Neoperla leopoldina Navás, 1932 , nomen dubium

Neoperla leopoldina Navás, 1932: 284 , fig. 83.

Navás named a specimen without abdomen from "Bomokandi (sources)". It was described as "similar to leroiana Klapálek. Larger ” [our translation]. Navás claimed that placement of the ocelli, shape of prothorax, colour of legs, and wing venation distinguished his new taxon “from species close to it”. The names of the allegedly related species were not disclosed. The characters in question are all insufficient for species identification. A doubtful taxon. We did not study the type (allegedly in MRAC) .

7. Acknowledgements

H. B. Noel Hynes was the first contemporary scientist dealing with the taxonomy of the African Neoperla . He recognised that the previously used habitus characters are unsuitable for species distinction and instead looked at genitalia and eggs. He concluded that the African fauna comprises

* either one single variable species of Neoperla .

* or many very similar and often coexisting ones ( Hynes 1952a).

We repeatedly discussed our controversial opinions on the subject, and Noel eventually considered there might be two, possibly even three different African Neoperla . I (PZ) would like to honour my late friend but realise that he would not have appreciated an African Neoperla species named after him.

The few SEM-micrographs in our paper were either prepared by technicians at the former Max-Planck-Institute of Limnology at Pl̂n while PZ worked in the institute's branch office at Schlitz. Others were adopted from T. Moore (1991), with permission of B. P. Stark ( Clinton , Mississippi, USA) to use illustrations from the work of his former student; we thank him very much .

We are particularly grateful to Hong Shen (SMNS at the time), Diana Hartley, James Nicholls and Stephen Bent (all CSIRO) for their technical support with the generation and assembly of DNA sequence data. The genome-skimming approach utilised developments and labour resources from AZ‘s ”High-throughput Collection Genomics“ project in CSIRO‘s Environomics Future Science Platform program.

We sincerely thank S. Grubbs for constructive suggestions and careful editing of our manuscript.

The present work was only possible because many individuals and several institutions supported us by loan or gift of specimens, help with literature, or advice. We are very grateful to all of them and list them in alphabetical order: J. S. Amakyé ( Ghana); Trond Andersen (Zoological Museum, University of Bergen, Norway); Richard W. Baumann (BYU, Provo, USA); Per Brinck † (Lund, Sweden); Daniel Bartsch (Stuttgart, Germany); Claude Berthélemy † (Toulouse, France); Klaus B̂ttger † (Wolfenbüttel, Germany); Alice-Marie Buset & Stéphane Hanot (MRAC, Tervuren, Belgium); Pavel Chvojka (Prague, Czech Republic); Carlo Consiglio (Roma, Italy); Samuel Danflous (Toulouse, France); Museum für Naturkunde, Berlin (MFNB; J. Deckert, W. Mey, M. Ohl, Germany); R. Edward Dewalt, Champaign, Illinois ( USA); Jean-Marc Élouard (Lauret, France); Oliver S. Flint † (USNM, Washington D. C., USA); Romolo Fochetti (Viterbo, Italy); Wolfram Graf (Vienna, Austria); C. Grinter, D. Kavanaugh (CAS, San Francisco, USA); Lars Hendrich (ZSM, München, Germany); B. Huber and H. Krammer (Museum Koenig, Bonn, Germany); Joachim Illies † (Schlitz, Germany); Vladimir Landa † (Ceské Budejovice, Czech Republic); Loic Matile † (MNHN, Paris, France); Ryuichiro Machida & Shodo Mtow (Univ. of Tsukuba, Japan); Michael Mühlenberg (Ĝttingen, Germany); Davíd Muranyi (Budapest and Eger, Hungary); Mathias Nuss (Dresden, Germany); Mike Picker (Cape Town, Rep. South Africa); Volker Puthz (Schlitz, Germany); Susanne Randolf (NHMW, Austria); G. de Rougemont † (Oxford, G.B.); Rafal Ruta (Wroclaw, Poland); Michel Sartori (Lausanne, Switzerland); D. Schimrosczyk (Warsaw, Poland); Peter Schüle (Herrenberg, Germany); Tomaš Soldán † (Ceské Budejovice, Czech Republic); F. Song ( China Agricultural University, Beijing); Bernhard Statzner † (Karlsruhe, Germany); David Stevens (Cape Town, Rep. South Africa); Manuel Tierno de Figueroa ( Granada, Spain); Wolfgang Tobias (Oberursel, Germany); Thomas Wagner (Koblenz, Germany); Peter Ward (BMNH, London, G.B.); Jason D. Weintraub (ANSP, Philadelphia, USA); Dr. Whitehead (South African Museum, Cape Town, Rep. South Africa).

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