Europharinodes Yin & Cai, 2022

Yin, Zi-Wei, Tihelka, Erik, Lozano-Fernandez, Jesus & Cai, Chen-Yang, 2022, The first fossil Hybocephalini (Coleoptera: Staphylinidae: Pselaphinae) from the middle Eocene of Europe and its evolutionary and biogeographic implications, Arthropod Systematics & amp; Phylogeny 80, pp. 279-294 : 279

publication ID

https://dx.doi.org/10.3897/asp.80.e82644

publication LSID

lsid:zoobank.org:pub:1D57CB0C-56C4-4DA5-91E1-CF7E9BB40E74

persistent identifier

https://treatment.plazi.org/id/0B631F8C-D490-4174-89F9-EE6274592F92

taxon LSID

lsid:zoobank.org:act:0B631F8C-D490-4174-89F9-EE6274592F92

treatment provided by

Arthropod Systematics & Phylogeny by Pensoft

scientific name

Europharinodes Yin & Cai
status

gen. nov.

Europharinodes Yin & Cai View in CoL gen. nov.

Figs 2 View Figure 2 , 3 View Figure 3 , 4 View Figure 4 , 5 View Figure 5

Type species.

Europharinodes schaufussi sp. nov.

Diagnosis.

Most of body covered with squamous setae; body generally compact; head, pronotum and femora coarsely punctate. Head roundly-triangular, with short and narrow frontal rostrum; large vertexal and frontal foveae present; antennal insertions close, antennomeres slightly elongate to sub-moniliform, clubs formed by apical three enlarged antennomeres; lacking ocular-mandibular carinae; clypeus sharply sloping, anterior margin carinate and rounded, lateral margins straight to eyes; maxillary palpi small, four-segmented, with short apical palpal cone. Pronotum with median and lateral antebasal foveae obscured by squamous setae; with antero-hypomeral foveae, lacking hypomeral carinae. Elytra each with two large basal foveae and two distinct discal striae. Each tarsus plesiomorphically with two subequal claws. Abdomen with tergite 1 (IV) slightly longer than 2 (V), tergites 1-4 (IV-VII) broadly sulcate at bases; paratergites moderately broad and laterally protruding. Aedeagus symmetrical, basal capsule enlarged, paired parameres elongate.

Description.

Habitus (Figs 2B-D View Figure 2 ; 3A, B View Figure 3 ) stout, compact; most of body with squamous setae. Head (Fig. 3E View Figure 3 ) roundly triangular; vertex coarsely punctate, vertexal foveae (= dorsal tentorial pits; Fig. 3E View Figure 3 ; vf) relatively large, obscured by squamous setae; frontal fovea (Fig. 3E View Figure 3 ; ff) distinct, rostrum short and narrow, antennal insertions located at ventral surface of rostrum, tubercles barely raised; compound eyes (Fig. 3G View Figure 3 ; ce) large, prominent; lacking ocular-mandibular carina; clypeus (Fig. 3G View Figure 3 ; cl) sharply sloping, with broadly rounded, ridged anterior margin; labrum (Fig. 3G View Figure 3 ; la) transverse, subtrapezoidal; mandible (Fig. 3G View Figure 3 ; ma) with single distinct apical and preapical teeth; maxillary palpus small (Fig. 3H, I View Figure 3 ), four-segmented, with palpomere 1 small, indistinct, 2 (Fig. 3H View Figure 3 ; p2) pedunculate basally and broadened toward apex, 3 (Fig. 3H View Figure 3 ; p3) roundly triangular, 4 (Fig. 3H View Figure 3 ; p4) sub-fusiform, with short palpal cone at apex. Venter with small, widely separated gular foveae (= posterior tentorial pits; Fig. 3F View Figure 3 ; gf) in shared transverse impression, gular moderately raised along middle, weakly impressed admesally; neck region broad. Paired tentorial arms (Fig. 3E View Figure 3 ; ta) V-shaped, each branched at base and extending anteriorly to reach inner wall of clypeus (right arm broken at apex in holotype). Pharynx (Fig. 3E View Figure 3 ; ph) about 2/3 of head length, posteriorly broadened. Antenna 11-segmented, moderately elongate, extending to approximately half of elytral length when extending posteriorly, antennomeres each slightly elongate to sub-moniliform, club (Fig. 3F View Figure 3 ) formed by enlarged apical three antennomeres.

Pronotum (Fig. 4A View Figure 4 ) approximately as long as broad, sides rounded, broadest at middle, anteriorly and posteriorly narrowed, surface with dense squamous setae (Fig. 3C View Figure 3 ); disc coarsely punctate, with squamous setae all over surface, covering median and lateral antebasal foveae (Fig. 4A, B View Figure 4 ; maf, laf), lacking sulcus or carina. Hypomeron extended, lacking carina, with anterior hypomeral fovea (Fig. 4B, C View Figure 4 ; ahf). Prosternum (Fig. 4C View Figure 4 ) with widely separated lateral procoxal foveae (Fig. 4C View Figure 4 ; lpcf), anterior part shorter than coxal part, margins of coxal cavity moderately carinate.

Elytra broadly truncate at bases, each elytron with two large basal foveae (Fig. 4D View Figure 4 ; bef), with two long discal striae (Fig. 4G View Figure 4 ; ds) extending from foveae to posterior elytral margin, lacking subhumeral fovea or marginal sulcus. Metathoracic (hind) wings (Fig. 2D View Figure 2 ; hw) fully developed.

Mesoventrite with median foveae (Fig. 4E View Figure 4 ; mmsf) originating from shared transverse impression, moderately separated, with pairs of large lateral and distinct anterolateral mesoventral foveae (not shown in figure); metaventrite with large, setose lateral coxal foveae (Fig. 4F View Figure 4 ; lmcf), with single median metaventral fovea (Fig. 4E View Figure 4 ; mmtf), posterior margin with small, narrow split at middle (Fig. 4E View Figure 4 ; ms); metacoxae broadly separated. Marginal carina of meso- and metaventrite complete (Fig. 4F View Figure 4 ; mc).

Legs (Fig. 5A-C View Figure 5 ) moderately elongate; femora roughly punctate; with short tarsomere 1 and long tarsomere 2 and 3, 2 slightly longer than 3, each tarsus with two subequal pretarsal claws (Fig. 2G View Figure 2 ).

Abdomen constricted at base, with dense squamous setae (Fig. 3D View Figure 3 ); tergite 1 (IV) slightly longer than 2 (V), 2-4 (V-VII) subequal in length, tergites 1-4 (Fig. 4G, H View Figure 4 ; t1-4) each deeply and broadly sulcate at base, at lease with one pair of basolateral foveae (Fig. 4G View Figure 4 ; blf), tergite 5 (VIII) (Fig. 4H View Figure 4 ; t5) roundly triangular, transverse, medially roundly emarginate at posterior margin; sternite 2-5 (IV-VII) (Fig. 4H, I View Figure 4 ; s2-5) each broadly sulcate at base, with one pair of basolateral foveae (Fig. 3H View Figure 3 ; blf), 6 (VIII), sternite 6 (Fig. 4H, I View Figure 4 ; s6) transverse.

Males have modified sternites 4 and 5 (VI and VII). Aedeagus (Fig. 5D-F, I-K View Figure 5 ) dorso-ventrally symmetrical.

Etymology.

The new generic name is a combination of Latin ' Eurōpa (Europe)' and genus Apharinodes , referring to the origin of the fossil in Baltic amber and its affinity with Apharinodes . The gender is feminine.

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Coleoptera

Family

Staphylinidae

SubFamily

Pselaphinae

Tribe

Hybocephalini