Steatonyssus spinosus Willmann, 1936

Orlova, M. V., Klimov, P. B., Orlov, O. L., Smirnov, D. G., Zhigalin, A. V., Budaeva, I. V., Emelyanova, A. A. & Anisimov, N. V., 2021, A checklist of bat-associated macronyssid mites (Acari: Gamasina: Macronyssidae) of Russia, with new host and geographical records, Zootaxa 4974 (3), pp. 537-564 : 553

Steatonyssus spinosus Willmann, 1936

Steatonyssus spinosus Willmann, 1936: 152 .

Steatonyssus spinosus .— Fonseca, 1948: 319; Yunker & Radovsky, 1966: 96; Beron, 1968: 160; 1969: 163; 1971: 100; Solomon, 1971: 313; Mahnert, 1971: 926; Pinchuk, 1971: 110; 1976: 129; Chow, 1973: 53; Rybin, 1983: 359; Uchikawa & Kumada, 1977: 427; Haitlinger & Ruprecht, 1977: 333; Haitlinger, 1978: 713; Ogadzhanyan & Arutunyan, 1974: 80; Teng, 1980; Rybin, 1983: 359; Schmidt, 1987; Stanjukovich, 1990: 197; Haitlinger & Lupicky, 2008: 124.

Steatonyssus (S.) spinosus .— Till & Evans, 1964: 567; Ah & Radovsky, 1967: 425; Radovsky 1966: 94; 1967: 207; 2010: 101; Beron 1973: 124; Orlova et al., 2011: 136.

Steatonyssus musculi .— Dusbábek, 1964b: 103.

Type locality: Zoo of Hamburg ( Germany) ( Willmann, 1936) .

Type host: [ Solenodon paradoxus Brandt, 1833 ] (from feces) ( Willmann, 1936).

Material. M ex Pipistrellus pygmaeus from biological station “Venevitinovo”, 4 June 2019, leg. I. V. Budaeva; F, 16 N1 ex Pipistrellus nathusii from State Nature Reserve “Samarskaya Luka”, 1-31 July 2006, leg. D.G. Smirnov; 2 N1 ex Vespertilio murinus from Biological station “Kolarovo”, 11 September 2015, leg. A. V. Zhigalin; N1 ex Myotis sibiricus from Tigirekskiy State Natural Reserve, 16 June 2016, leg. A. V. Zhigalin, O.L. Orlov; N1 ex Myotis sibiricus from State Natural Reserve “Barguzinskiy”, 2 July 2015, leg. D. V. Kazakov; N1 ex Myotis petax from Okhotnichiya cave, 4 September 2015, leg. D. V. Kazakov; F, M, N1 ex Eptesicus nilssonii from lake Gusinoe, 30 June 2016, leg. D. V. Kazakov; N1 ex Myotis sibiricus from Argaley-3 cave, 21 May 2016, leg. D. V. Kazakov; M ex Ep. nilssonii from Nizhniy Tsasuchey, 18 July 2015; M, 2 N1 ex Myotis davidii from Adun-Chelon, 24 July 2015, leg. D. V. Kazakov.

Distribution in Russian Federation: Samara Province (present paper), Volgograd Province ( Stanyukovich, 1997), Bashkiria Republic (present paper), Sverdlovsk Province (Orlova, 2013; Orlova et al., 2017c), Chelyabinsk Province (Orlova, 2013; Orlova et al., 2017c), Novosibirsk Province ( Orlova et al., 2015d), Altai Region (present paper), Tomsk Province ( Orlova et al., 2017c), Tuva Republic (Orlova et al., 2015a), Irkutsk Province (present paper), Buryatiya Republic (present paper), Zabaykalskiy Region ( Orlova et al., 2017c), Primorskiy Region ( Medvedev et al., 1991), [Ural] ( Orlova et al., 2015d), [Southern Siberia] ( Orlova et al., 2015c), [Far East] ( Stanyukovich, 1997).

Distribution outside Russian Federation: Europe ( Radovsky, 1967). Asia ( Radovsky, 1967).

Hosts. [ Solenodon paradoxus ], Vespertilio murinus ( Stanyukovich, 1997; Orlova et al., 2015d; present paper), V. sinensis (as V. superans — Till & Evans, 1964; Medvedev et al., 1991), Myotis myotis ( Stanyukovich, 1997) , M. blythii ( Stanyukovich, 1997) , M. dasycneme ( Orlova et al., 2015d) , M. daubentonii ( Stanyukovich, 1997) , M. mystacinus ( Stanyukovich, 1997) , M. petax ( Orlova et al., 2015c; present paper), M. davidii (present paper), M. sibiricus (present paper), Hypsugo alashanicus Bobrinskoy, 1926 (as Pipistrellus savii — Medvedev et al., 1991), Pipistrellus pygmaeus (present paper), Pi. pipistrellus ( Stanyukovich, 1997) , Pi. nathusii ( Orlova et al., 2015d; present paper), Eptesicus nilssonii ( Orlova et al., 2015d; present paper), Ep. serotinus ( Stanyukovich, 1997) , Nyctalus noctula ( Stanyukovich, 1997) , Ny. leisleri ( Stanyukovich, 1997) , Ny. aviator ( Beron, 2014) , Barbastella barbastellus ( Stanyukovich, 1997) , Plecotus auritus ( Stanyukovich, 1997) , Pl. ognevi (present paper), Rhinolophus ferrumequinum ( Stanyukovich, 1997) , Rh. hipposideros ( Stanyukovich, 1997) .