Pancheria lanceolata (Pamp.) Baker

Hopkins, Fortune & Bradford, Jason C., 2009, Nomenclature and typification of names in the endemic genus Pancheria (Cunoniaceae) from New Caledonia, Adansonia (3) 31 (1), pp. 103-135 : 120-121

22. Pancheria lanceolata (Pamp.) Baker f.

Journal of the Linnean Society, Botany 45: 302 (1921) , with indirect reference to the basionym: Pancheria alaternoides Brongn. & Gris var. lanceolata Pamp., Annali di Botanica (Rome) 2: 95 (1905). — Types (as given in protologue of basionym): “In collibus circa Wagap [Vieillard, – Herb. de la Nouvelle Calédonie, 1861-67, n. 598, n. 581?] – specimen foem. – (B-B, DC)”. — Lectotype (here designated): Wagap, [1861-1867], old fr., Vieillard 598 = 581? (G! ex DC; isolecto-, G! ex B-B; K!; P! × 3, P00143085 , P00602423 fragm. “A”, P00602420 fragm. “A”).

REMARKS

Vieillard wrote the name Pancheria lanceolata on specimens but never published it. Most of the specimens at P with this name either in Vieillard’s hand or copied onto his original labels are now identified as P. gatopensis (e.g., Vieillard 2064, Wagap, various dates, old fr. or ♂ fl.; Deplanche 2064 (Vieill.) and/or Mus . Neocal. 62, Pum(e), various dates, old fr. or ♂ fl.) while Pancher 46481, “Cougui” (= Koghi) is a mixture of P. gatopensis and P. ferruginea .

Pampanini (1905) published the name Pancheria alaternoides var. lanceolata without reference to Vieillard’s unpublished name and as neither of the two syntypes at G has “ P. lanceolata Vieill. mss” marked on them, the epithet lanceolata must be ascribed to Pampanini alone.

Guillaumin (1911a: 135) invalidly published the name “ P. lanceolata Vieill. mss.” as a nomen nudum in synonymy under P. alaternoides var. lanceolata Pamp.

Baker (1921: 302) published the name “ P. lanceolata Vieill. ” without reference to Pampanini or Guillaumin, and mentioned Compton 2382, as belonging to this taxon. This can be considered an indirect reference to P. alaternoides var. lanceolata Pamp. , through Guillaumin’s publication (1911a: 135), and hence Baker made a new combination, and the type of P. lanceolata is the same as for P. alaternoides var. lanceolata . However, Compton 2382 is now identified as P. gatopensis .

Däniker (1932:170) published the name “ Pancheria lanceolata Pampanini [...] 1905” and mentioned “ Pancheria lanceolata Vieill. mss.; Baker f. in Journ. of Linn. Soc. XLV (1921) 302” in synonymy.

Guillaumin (1941: 249) stated that Pancheria alaternoides var. lanceolata Pamp. did not correspond to P. lanceolata Vieill. mss., in contrast to his opinion in Guillaumin (1911a), and on p. 250, he gave the latter name as a synonym of his new species P. gatopensis . On p. 252 he mentioned two specimens that were identical to P. lanceolata but without mentioning an author. Guillaumin (1948) made no mention of either P. lanceolata or P. alaternoides var. lanceolata .

Although the lectotype of Pancheria alaternoides var. lanceolata , the basionym of P. lanceolata , appears to have two collection numbers, they refer to a single gathering and indicate that Vieillard thought his species 598 was possibly the same as his species number 581. However, three separate gatherings have the number 598. First, Vieillard 598 = 581? (Wagap, 1861-1867, P!), the lectotype of P. alaternoides var. lanceolata , has simple, shortly petiolate, lanceolate leaves (“A”). Second, Vieillard 598 (Kanala, 1855-1860) is the type of P. alaternoides ( var. alaternoides ), which is a separate, well-defined taxon with simple, almost sessile, elliptic leaves. Finally, a sheet at P labelled “ Vieillard 598, Kanala etc, 1861-67” was determined (by Vieillard?) as P. alaternoides but it is not; it has a mixture of simple leaves below and distinctly petiolate compound leaves above and is P. cf. reticulata (“B”). Two other sheets at P are mixtures of “A” and “B”.

Although the type of Pancheria lanceolata was originally described as a variety of P. alaternoides , it is only superficially similar to that species since the former has narrower leaves with a distinct petiole and more of the margin is toothed than is typical in P. alaternoides . The type of P. lanceolata also has some similarities to P. gatopensis but in the latter the leaves are considerably broader, the peduncles much shorter, and the young stems, leaves and peduncles have dense indumentum. Pancheria lanceolata is possibly a hybrid, and if so, its parentage is likely to involve P. gatopensis and P. elegans , rather than P. alaternoides . Pancheria alaternoides is confined to the extreme south of Grande Terre, and even allowing for the lack of reliability and precision of Vieillard’s locality data, the type of P. lanceolata appears to have come from further north, since Wagap is near the estuary of the Tiwaka river in northeastern Grande Terre. The few recent gatherings that are morphologically close to the type of P. lanceolata (Pillon & Barrabé 331, Cap Bocage; Thorne 28057, Tiebaghi; Thorne 28289, Col de Ho) are also from outside the geographical range of P. alaternoides .

Pancheria lanceolata can also be compared with material labelled by Guillaumin as “ P. alaternoides var. angustifolia ” (nom. inval.), but the leaves of P. lanceolata are larger, more ovate-elliptic, and have longer petioles. The two are not sympatric and specimens in the latter group are all from the south of Grande Terre.