Ononis tridentata var. edentula Webb ex Willkomm (1877: 413)

Ferrer-Gallego, P. Pablo, 2023, Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae), Phytotaxa 598 (1), pp. 32-58 : 52-55

Ononis tridentata var. edentula Webb ex Willkomm (1877: 413)

Willkomm’s protologue of Ononis tridentata α [var.] edentula includes a brief diagnosis “foliis lineari-lanceolatis linearibusve canaliculatis, interdum subteretibus, integerrimis mucronatis”, preceded by the epithet “α. edentula Wbb.” and followed by these two names and indications: “Cicer silv. fruticans, angustifolium non serratum, Hispanicum Barr. Ic. 300! O. tridentata Willk. Sert. p. 41 et pl. exs. 1850, n. 530, Bourg. pl. exs. n. 1724!. In the protologue also was mentioned several gatherings: “[…] in Catalaun. int. Hortalets et Tost atque in Conca de Tremp, CSTA.!, in regno Valent. pr. Játiva, BOURG.! in Serranía de Cuenca int. Mira et Villora, WK., int. urbem Jaen et Campillos de Arenas, WK. […]” (see Willkomm & Lange 1877: 413).

I have found several syntypes at COI and P herbaria. Willkomm cited in the protologue material collected in Cuenca: “in Serranía de Cuenca int. Mira et Villora, WK”. A specimen belonging to this gathering is kept at COI, with barcode COI00053673. This material has been selected as the lectotype of the name O. tridentata f. angustifolia (see above) and is also a syntype of the name O. tridentata var. edentula .

On the other hand, there is a specimen at COI, with barcode COI00053680 that is part of the Bourgeau’s exsiccatum cited in the protologue as “Bourg. pl. exs. n. 1724” and also as “in regno Valent. pr. Játiva, BOURG.!”. This specimen are three well-preserved stems, with leaves and flowers, and an original printed label of the Bourgeau’s exsiccatum, annotated as: “E. Bourgeau, Pl. d’Espagne, 1852 / 1724. Ononis tridentata, Ass. / ver. edentula Webb. / (Webb.) / Collines arides près S. Felipe de Xativa. / 17 Juin.” ( Fig. 13 ). In the herbarium P there are several duplicates of Bourgeau 1724, with barcodes P03600614, P03447655, P03447761, P03447763, P03447771. These specimens are syntypes.

In the same sheet that the specimen COI00053680 is preserved the specimen barcoded COI00053681. This specimen consists of two stems, with leaves and flowers, and is accompanied by an original handwritten label by Willkomm ( Fig. 13 ). The material was collected on August 22, 1845 from the locality cited in the protologue as “int. urbem Jaen et Campillos de Arenas, WK.”, this material is therefore also a syntype.

In conclusion, among the original material, I designate as the lectotype of Ononis tridentata var. edentula the specimen COI00053680 from the Bourgeau’s exsiccatum. The lectotype selected is a complete and well-preserved specimen that shows relevant diagnostic features (e.g., leaflets blade linear to linear-ensiform, more than six times longer than wider, glabrous, with 0–1(2) apical teeth), and it clearly represents the traditional concept of this name ( Willkomm & Lange 1877) and reflects the current application of Ononis tridentata var. edentula as a synonym of Ononis tridentata subsp. angustifolia (see e.g., Devesa & López González 1997, Devesa 2000, López González 2001).

Figures

FIGURE 13. Lectotype of Ononis tridentata var. integrata Amo, COI (barcode COI00053681) and lectotype of Ononis tridentata var. edentula Webb ex Willkomm, COI (barcode COI00053680). Image courtesy of the herbarium COI, reproduced with permission.