Ononis tridentata var. canescens Lamarck ex Candolle (1825: 161)

Ferrer-Gallego, P. Pablo, 2023, Original elemets and typification of the names included in the Ononis tridentata complex (Fabaceae), Phytotaxa 598 (1), pp. 32-58 : 40-42

Ononis tridentata var. canescens Lamarck ex Candolle (1825: 161)

Ononis tridentata var. canescens was described by Candolle (1825: 161) with the accompanying diagnosis “ramis cano-tomentosis, foliis pallidis cuneiformibus quinquedentatis”, and followed by the provenance “in Arragoniâ” and the references “Asso fl. arr. n. 671. var. 2” from Asso (1779: 95 [2. Varietas]) and “Barr. icon. t. 416 [see 419]” from Barrelier (1714: ic. 419 [not 416]). The protologue also includes a reference to Lamarck’s Encyclopédie méthodique (1785: 508), as “ β. canescens (Lam! dict. I. p. 508)”. The indication in the protologue as “(v. s.)” must be treated as the acronym of “ vidit sicco ” (i.e., seen in a herbarium sheet).

Lamarck (1785: 508) mentioned as a variety into Ononis tridentata or his “Bugrane à trois dents”: “ β. Eadem? foliis cuneiformibus quinquedentatis, pedunculis unifloris. Flor. Aragon. p. 95. nº. 671. var. 2. Cicer montanum fruticans, incanum, angustifolium, serratum minus. Barrel. Ic. 419” and “La plante β forme pareillement un sous-arbrisseau trèsrameux & haut d’un pied ou d’un pied & demi; mais elle en diffère par plusieurs caractères assez remaquables. Ses tiges sont blanches & beaucoup plus cotonneuses; ses feuilles sont plus courtes, cuneiforms, blanchâtres, & la plupart à cinq petites dents à leur sommet. Les pédoncules sont courts, latéraux, plus souvent uniflores, & soutiennent des fleurs purpurines dont le calice est moins velu. Ces plantes croissant en Espagne, & nous ont été communiquées par M. l’Abbé Pourret (v. s.)”.

The reference of Barrelier (1714: t. 419) provided an illustration that can be considered original material used by Candolle to describe O. tridentata var. canescens . On the other hand, in the Lamarck’ herbarium at P, there is a sheet, with barcode P00296101, which bears four stems, with leaves, flowers, and fruits, and an original label handwritten by Pourret and Lamarck, annotated as “ Ononis tridentata / varietas pedunculis, floris / calycibus glabris / ad Balbastr. / in Arrag. / [handwritten by Pourret?] Barrel. ic. 419 [handwritten by Lamarck]” (image available at https://mediaphoto. mnhn.fr/media/1665502967319P49a40tO4W7dyGh7). This specimen matches the description published by Lamarck (1785: 508) and Candolle (1825: 161). In addition, there is a sheet in the Candolle herbarium at G-DC, with barcode G00477895 ( Fig. 6 ), this specimen can be treated as a duplicate of the sheet at P (P00296101). The sheet preserved at G-DC bears two stems, with leaves and fruits, and two original labels handwritten by Candolle, annotated as “ Arragon ” and “ Ononis tridentate β. Lam.” ( Fig. 6 ).

In conclusion, among the original elements, the illustration published by Barrelier (1714: ic. 419 [not 416 as was cited in the protologue]) and the specimen at G-DC, I designate as the lectotype of the name the specimen G00477895 .

Figures

FIGURE 6. Lectotype of Ononis tridentata var. canescens Lamarck ex Candolle, G (barcode G00477895). Image courtesy of the herbarium G, reproduced with permission.