Satsuma albida, (H. ADAMS, 1870)

Wu, Shu-Ping, Hwang, Chung-Chi & Lin, Yao-Sung, 2008, Systematic revision of the arboreal snail Satsuma albida species complex (Mollusca: Camaenidae) with descriptions of 14 new species from Taiwan, Zoological Journal of the Linnean Society 154 (3), pp. 437-493 : 442-444

SATSUMA ALBIDA (H. ADAMS, 1870) ( FIGS 3 , 4 )

Helix (Satsuma) albida H. Adams, 1870: 378 , pl. 27, fig. 9. (Nov., 1870)

Helix albida, Pfeiffer, 1876: 379–380 ; Moellendorff, 1884: 335–336; Tryon, 1887: 218, pl. 51, fig. 53.

Helix (Satsuma) albida, Schmacker & Boettger, 1891: 159 .

Ganesella albida, Pilsbry, 1895: 169 ; Pilsbry & Hirase, 1906 [1905]: 736; Kuroda, 1941: 144; Kuroda, 1958: 145.

Satsuma albida, Minato, 1976: 84 ; Richardson, 1985: 268; Chang et al., 1996: 25–30 [non albida ]; Hsieh et al., 2006: 231.

Coniglobus (Luchuhadra) albida, Chang, 1984: 15 , pl. 5, fig. 28.

Satsuma albidum, Lai, 1990: 49 . [incorrect gender ending]

Satsuma (Coniglobus) albida, Ohara & Otani, 2002: 83 .

Material examined

Type specimen: Only one specimen originally collected by R. Swinhoe was found and is here desig nated as the lectotype ( BMNH 1878.1.28.229, SH, shell height; SD, shell diameter; SH/SD, shell height/shell diameter ratio; Whorls, number of whorls; N, sample size of all type specimens for new species or type(s) and additional specimens measured in this study for the three already described species, S. albida , S. insignis and S. mollicula ; median (bold), minimum and maximum (parentheses).

Fig. 3A–D ), which was part of H. Adams’ collection purchased by the Natural History Museum, London. The number of specimens was not mentioned in Adams’ description, but a later mark on the case, ‘paratype’, indicated that there was probably more than one specimen examined by H. Adams although no specimen and designation of types can be traced. Therefore, we prefer to designate the single specimen as lectotype.

Additional material: Nineteen specimens were collected from Nanhua (N = 1), Jiaxian (N = 2), Baolai (N = 5), Shanping (N = 1) and Dajin (N = 10). Three specimens from Dajin were deposited as vouchers,

TMMT P- 0277 (dissected, in alcohol), BMNH 20060782 (shell) and ANSP 413693 (shell).

Type locality

Taiwan, Formosa. The locality ‘Taiwan’ indicated the ‘ Taiwan Hsien (County), Taiwan Fu’ on south-west Formosa at that time ( Swinhoe, 1864), which is equivalent approximately to part of Tainan County and Kaohsiung County at present. The label accompanying with the specimen is marked as ‘Takow, Formosa’ and is incongruent with H. Adams’ description. Takow (or Takao) located at Qijin, coastal part of Kaohsiung City nowadays, is less likely to have this species. We suggest that it is either mislabelled by

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1 03 0 5 07 8 0 (00) 0 (0) 0 0 0

1 0 0 0 0 0 7 0 1 0 0 0 0 6 7 0 0 0 (0 0) 0 0 0 0 1 0 0 0 5 0 7 8 1 0 0 4 0 0 7 0 1 0 0 4 0 0 7 8

1 0 0 0 5 6 7 8 1 0 0 0 (5 6 7) 8

0 0 0 0 0 6 0 0 0 0 0 0 0 0 7 0 1 0 0 0 0 6 7 8 1 0 0 0 0 0 7 8

1 (0 0 0) 5 0 7 0 1 0 (0 4) 5 0 7 8 1 0 3 0 5 0 7 0 0 0 0 0 0 0 0 0 1 0 0 4 0 6 7 8 0 0 0 0 0 6 7 0

Figures

Figure 3. Satsuma albida. A–D, lectotype (BMNH 1878.1.28.229, shell height = 15 mm). E, living specimen. Arrow indicates the pre-apertural constriction.

Figure 4. Reproductive system of Satsuma albida (TMMT P-0277, Dajin). A, whole genitalia; B, interior of genitalia. Scale bar = 5 mm. See text for abbreviations.