Novalena clara, Maya-Morales, Julieta & Jiménez, María Luisa, 2017

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 39-40

Novalena clara new species

Figures 1 d, h, 19; Map 4

Type. Holotype male: MEXICO: Chihuahua: Sierra del Nido, Arroyo del Álamo , 29°20′N, 106°50′W, 27.XII.1969, V. Roth (CASENT 9048865), deposited at CAS .

Etymology. The specific name is a noun in apposition referring to the color tone of the holotype ‘clara’ which means ‘ light’.

Diagnosis. The males of this species differ from N. approximata and N. bosencheve in having the tegular median process with one projection ( Fig. 19 a) and the RTA with a denticle on ventral projection ( Fig. 19 b). The females differ from N. approximata in having the spurs more conspicuous ( Fig. 19 c), from N. franckei and N. volcanes in having the primary spermathecae with posterior part wider than anterior part ( Fig. 19 d), from N. paricutin in having the spurs directed mesally, from N. saltoensis in having the spurs separated from the posterior margin by their basal width, and from N. creel in having the spurs in lateral position ( Fig. 19 c).

Description. Male (holotype): Carapace yellow. Chelicerae yellow, condyles yellow. Endites yellow. Labium light brown with white tip. Sternum brown with a central yellow band with arrowhead shaped spot. Legs yellow with black spots on coxa and trochanter, three black rings on femur, two on patella, and two on tibia. Opisthosoma light brown with an anterior reddish spot and one yellow dorsal band. PLS with basal segment brown. Total length 10.13. Carapace length 4.5, width 3.13, cephalic region width 1.88, ocular region width 1. Eye diameter: AME 0.13, ALE 0.19, PME and PLE 0.17. Distance between eyes: AME-AME 0.15, AME-ALE 0.12, AME-PME 0.1, ALE-PLE 0.1, ALE-ALE 0.5, PME-PME 0.19, PME-PLE 0.19. Clypeus height 0.52. Chelicerae with four retromarginal teeth; basal segment length 1.33, fang length 0.76. Labium as long as wide (0.61/0.64). Endites slightly convergent (distance at their base compared at their tips 0.64/0.42). Sternum longer than wide (2.24/1.9). Opisthosoma longer than wide (5.88/3.38). ALS separated by less their basal diameter (0.19/0.48), PLS with distal segment shorter than basal segment (0.43/0.71). Leg lengths: I—femur 3.5/ patella-tibia 4.5/ metatarsus 3.38/ tarsus 2.38; II—3.75/ 4.5/ 3.5/ 2.13; III—3.75/ 4.38/ 3.5/ 1.88; IV—4.38/ 5.13/ 5/ 2.38. Spination: femur dorsal I— 2-1 -2/ II—2-1 -3/ III—2-1 -3/ IV—2-1 -2; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 1-1-0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—1-1 -0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0; II—0-0-2/ 2-2-2/ 1-1-1/ 0-0-1; III—2-1 -2/ 2-2-2/ 0-1-1/ 1-0-1; IV—2-1 -2/ 2-1-2-2/ 0-1-1/ 0-1-1. Number of trichobothria on tarsus: I—8, II—7, III—5, IV—7. Number of dorsal spines on pedipalp: femur 3, tibia 2. Cymbium length 1.76, width 1.12. Embolus straight, conductor distally notched; tegular median process with two broad and overlapping tips; dorsal projection of RTA longer than ventral projection, which presents a denticle ( Fig. 19 a, b).

Female (paratype) (CASENT 9048874): Carapace yellow, white feathery scales on cephalic region. Chelicerae reddish, condyles orange. Endites light brown. Labium brown with white tip. Sternum brown with a median yellow band with arrowhead shaped spot directed posteriorly. Legs yellow, tibia to tarsus reddish. Three black rings on femur and two on patella. Opisthosoma yellow with several lateral and ventral black spots and yellow foliage. Spinnerets yellow. Total length 11.63. Carapace length 5.63, width 3.75, cephalic region width 2.38, ocular region width 1.25. Eye diameter: AME, PME and PLE 0.21, ALE 0.25. Distance between eyes: AME-AME 0.12, AME- ALE 0.13, AME-PME 0.25, ALE-PLE 0.1, ALE-ALE 0.58, PME-PME 0.15, PME-PLE 0.23. Clypeus height 0.38. Chelicerae with four retromarginal teeth; basal segment length 1.29, fang length 0.95. Labium wider than long (0.95/0.76). Endites slightly convergent (distance at their base compared at their tips 0.95/0.48). Sternum longer than wide (2.63/2.25). Opisthosoma longer than wide (6.38/3.75). ALS separated by half their basal diameter (0.3/ 0.61), PLS with distal segment as long as their basal segment (0.64/0.64). Leg lengths: I—femur 4/ patella-tibia 4.5/ metatarsus 3.63/ tarsus 1.88; II—4 / 4.38/ 2.5/ 1.88; III—3.75/ 4/ 5/ 1.88; IV—4.63/ 5.63/ 4.75/ 1.5. Spination: femur dorsal I—1-2 -2/ II—1-2 -2/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 1-1-0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—0-1-0/ 2- 2-2/ 1-1-0/ 0; III—1-2 -1-2/ 2-2-2/ 0/ 0; IV—1-2 -1-2/ 2-2-2/ 0/ 0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0; II—0/ 2-2-2/ 0- 1-1/ 0; III—2-2 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—1-1 -2/ 1-1-1-1-1-2/ 1-1-1/ 0-0-1. Number of trichobothria on tarsus: I—8, II—7, III—9, IV—6. Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (1.58/0.91); atrium superficial; spurs lateral ( Fig. 19 c, e). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent ( Fig. 19 d).

Variation. Males (n=2): body length 10.13–10.75; carapace length 4.5–5.58; patella-tibia I length 4.5–5. Additional material examined. Paratypes: MEXICO : Chihuahua: same data as holotype, 1 ♂ (CASENT 9048864) ; same data except 14.X.1969, 1 ♀ (CASENT 9048874).

Distribution. MEXICO: Chihuahua (Map 4).

Figures

FIGURE 1. Novalena spp., habitus: a, e, N. approximata (Gertsch & Ivie 1936), female; b, f, N. alvarezi sp. nov., female; c, g, N. chamberlini sp. nov., male; d, h, N. clara sp. nov., female; i, m, N. dentata sp. nov., male; j, n, N. irazu sp. nov., female; k, o, N. iviei sp. nov., male; l, p, N. leonensis sp. nov., female; q, u, N. mexiquensis sp. nov., male; r, v, N. poncei sp. nov., female; s, w, N. puebla sp. nov., female; t, x, N. rothi sp. nov., female. a – d, i – l, q – t, dorsal view; e – h, m – p, v – x, ventral view. Scale bars: a, d, j, q, s, t = 4 mm; b, i, l, r = 3 mm; c, k = 2 mm.

FIGURE 19. Novalena clara sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm.