Novalena franckei, Maya-Morales, Julieta & Jiménez, María Luisa, 2017

Maya-Morales, Julieta & Jiménez, María Luisa, 2017, Revision of the funnel-web spider genus Novalena (Araneae: Agelenidae), Zootaxa 4262 (1), pp. 1-88 : 48-49

Novalena franckei new species

Figure 25 ; Map 7

Type. Holotype male: MEXICO: Guanajuato: Municipality of Acámbaro , Sierra de los Agustinos, 20.X.2006, A.F. Quijano, J. Ponce, and M. Villaseñor (CNAN-T1123), deposited at CNAN .

Etymology. The specific name is a patronym in honor of Oscar F. Francke, one of the collectors of the paratype, for his contribution to Arachnology in Mexico.

Diagnosis. The males of N. franckei differ from N. approximata and N. bosencheve in having the tegular median process with one projection, and from N. clara and N. creel in having the tegular median process sclerotized ectally ( Fig. 25 a). The females differ from N. clara and N. volcanes in having the atrium occupying less than half of plate width ( Fig. 25 c).

Description. Male (holotype): Carapace brown, white feathery scales on cephalic region. Chelicerae and condyles dark brown. Endites and labium dark brown with white tips. Sternum dark brown. Legs yellow with three rings on femur, one on patella, two on tibia, and two on metatarsus. Opisthosoma dark brown with posterior light brown foliage. Spinnerets orange. Total length 6.5. Carapace length 3.25, width 2.13, cephalic region width 1.19, ocular region width 0.76. Eye diameter: AME 0.15, ALE 0.19, PME and PLE 0.15. Distance between eyes: AME- AME 0.06, AME-ALE 0.06, AME-PME 0.17, ALE-PLE 0.04, ALE-ALE 0.35, PME-PME 0.08, PME-PLE 0.1. Clypeus height 0.15. Chelicerae with three retromarginal teeth; basal segment length 1.18, fang length 0.45. Labium wider than long (0.45/0.36). Endites almost straight (distance at their base compared at their tips 0.45/ 0.24). Sternum longer than wide (1.61/1.31). Opisthosoma longer than wide (3.25/2). ALS separated by less their basal diameter (0.19/0.29), PLS with distal segment shorter than basal segment (0.31/0.35). Leg lengths: I—femur 2.43/ patella-tibia 2.9/ metatarsus 2.2/ tarsus 1.67; II—2.33/ 2.86/ 2.19/ 1.57; III—2.38/ 2.86/ 2.19/ 1.43; IV—2.86/ 3.38/ 3.33/ 1.67. Spination: femur dorsal I—1-2 -2/ II—1-3 -3/ III—1-3 -3/ IV—1-2 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 0/ ventral 2-2-2/ prolateral 1- 1-0/ retrolateral 0; II—0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 1-2-2/ 1-1-0/ 0; IV—1-1 -0/ 1-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-0-1; II—0-1-2/ 2-2-2/ 0-1-1/ 0-1-1; III—3-1 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-1 -2/ 2-2-2/ 0-1-1/ 0-1-1. Number of trichobothria on tarsus: I—7, II—6, III—6, IV—6. Number of dorsal spines on pedipalp: femur 3, tibia 3. Cymbium length 1.18, width 0.67. Embolus slightly curved; conductor with ventral projection slightly longer than dorsal one; tegular median process with ectal margin sclerotized; dorsal projection of RTA larger and less sclerotized than ventral projection ( Fig. 25 a, b).

Female (paratype) (CNAN-T0952): Carapace yellow, ocular region black. Chelicerae and condyles dark brown. Endites and labium dark brown with white tips. Sternum dark brown. Legs yellow, coxa with diffuse black spot, tibia and metatarsus light brown. Three black rings on femur, one on patella, two on tibia, and two on metatarsus. Opisthosoma light brown with anterior reddish spot, five pairs of lateral spots, and foliage. Spinnerets light brown. Total length 9.13. Carapace length 3.88, width 2.75, cephalic region width 1.67, ocular region width 0.95. Eye diameter: AME 0.17, ALE and PLE 0.21, PME 0.19. Distance between eyes: AME-AME 0.1, AME- ALE 0.1, AME-PME 0.19, ALE-PLE 0.06, ALE-ALE 0.44, PME-PME 0.12, PME-PLE 0.17. Clypeus height 0.23. Chelicerae with three retromarginal teeth; basal segment length 1.52, fang length 0.62. Labium slightly wider than long (0.61/0.52). Endites almost straight (distance at their base compared at their tips 0.61/0.39). Sternum longer than wide (1.9/1.57). Opisthosoma longer than wide (5.5/3.5). ALS separated by half their basal diameter (0.19/ 0.38); PLS with distal segment shorter than basal segment (0.35/0.42). Leg lengths: I—femur 2.88/ patella-tibia 3.75/ metatarsus 2.38/ tarsus 1.75; II—2.88/ 3.38/ 2.25/ 1.75; III—2.75/ 3.25/ 2.63/ 1.5; IV—3.38/ 4.25/ 3.38/ 1.75. Spination: femur dorsal I—1-1 -2/ II—1-3 -3/ III—1-3 -3/ IV—1-1 -3; patella I—dorsal 1-1/ prolateral 1/ retrolateral 0; II—1-1 / 1/ 0; III—1-1 / 1/ 1; IV—1-1 / 1/ 1; tibia I—dorsal 1-0-0/ ventral 2-2-2/ prolateral 1-1-0/ retrolateral 0; II—1 -0-0/ 2-2-2/ 1-1-0/ 0; III—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; IV—1-1 -0/ 2-2-2/ 1-1-0/ 1-1-0; metatarsus I—0/ 2-2-2/ 0-1-1/ 0-0-1; II—0/ 2-2-2/ 0-1-1/ 0-0-1; III—2-2 -2/ 2-2-2/ 0-1-1/ 0-1-1; IV—3-2 -2/ 2-2-2/ 0-1-1/ 0-0-1. Number of trichobothria: I—6, II—7, III—6, IV—7. Dorsal spines on pedipalp femur: 3. Prolateral spines on tibia: 1-2. Epigynal plate wider than long (1.24/0.7); atrium superficial and occupying less than half the plate width; anterior spurs separated by more their basal width ( Fig. 25 c, e). Internal genitalia: copulatory ducts straight, primary spermathecae adjacent with a dorsal depression ( Fig. 25 d).

Additional material examined. Paratype: MEXICO : Guanajuato: same locality as holotype, 20°12′27.9″N, 100°41′22.2″W, 2450 m, 25.III.2006, O. Francke, A. Valdez, H . Montaño, C. Santibáñez, and G. Villegas, 1 ♀ (CNAN-T0952).

Natural history. The species is sympatric with N. dentata and N. poncei .

Distribution. MEXICO: Guanajuato (Map 7).

Figures

FIGURE 25. Novalena franckei sp. nov.: a, male pedipalp, ventral view; b, same, retrolateral view; c, epigynum, ventral view; d, internal genitalia, dorsal view; e, epigynum, posterior view. Abbreviations: A, atrium; C, conductor; CD, copulatory ducts; E, embolus; FD, fertilization ducts; MA, median apophysis; RTAd, distal projection of RTA; RTAo, dorsal projection of RTA; RTAv, ventral projection of RTA; S, spurs; S 1, primary spermathecae; S 2, secondary spermathecae; T, tegulum; TMP, tegular median process. Scale bars: a = 0.5 mm; e = 0.25 mm.