Ischnothyreus bifidus, Edward & Harvey, 2014

Edward, Karen L. & Harvey, Mark S., 2014, Australian Goblin Spiders Of The Genus Ischnothyreus (Araneae, Oonopidae), Bulletin of the American Museum of Natural History 2014 (389), pp. 1-144: 41-48

publication ID

http://doi.org/ 10.1206/865.1

persistent identifier

http://treatment.plazi.org/id/E466E016-FF82-FF8D-FF3B-7C7A3D9FFDF3

treatment provided by

Felipe

scientific name

Ischnothyreus bifidus
status

new species

Ischnothyreus bifidus   , new species Figures 17–22 View Fig View Fig View Fig View Fig View Fig View Fig , map 3

TYPES: AUSTRALIA: Queensland: Male holotype, female allotype, and 1 female paratype from rainforest leaf litter in Mount Lewis Forest Reserve, 16 km from highway, along Mount Lewis Road , 16.57611 ° S, 145.26416 ° E (2 Apr. 2009, K. Edward and J. Waldock) deposited in QM (³ holotype: QM S95923 View Materials , PBI _ OON 00005674; ♀ allotype: QM S95924 View Materials , PBI _ OON 00005675; ♀ paratype: PBI _ OON 5640) GoogleMaps   .

ETYMOLOGY: The specific epithet is of the Latin bifidus   , meaning ‘‘bifurcated’’ ( Brown, 1956), and refers to the distinctive bifurcated process on the anterior face of the male chelicerae of this species.

DIAGNOSIS: This species is closely related to I. hoplophorus   , sp. nov., as both males possess a clypeus that is concave medially (figs. 17H, 20A, B, 23G), the anterior face of the chelicerae is modified with complex processes (figs. 19A, B, 22C, 24C), and the hyaline grooves, anterolateral corners of the carapace, endites, and the lateral edges of the labium are all heavily sclerotized (fig. 17G–I, 23G–H). Females share a unique epigynal region that consists of a heavily sclerotized platelike structure anterior to an ovoid shaped epigynal atrium (17J, 23I). Ischnothyreus bifidus   , sp. nov., can be distinguished by the bifurcated anterior process on the male chelicerae (fig. 22C), and the stouter platelike sclerite of the female epigynal region, which has straighter edges (figs. 21D, 22E).

MALE (PBI_OON_0005674, figs. 17A–C, G–I, 22A–D; paratype ³: figs. 18–19, 20A– C, E, 21A–C). Total length 1.81. CEPHA- LOTHORAX: Carapace brown, broadly oval in dorsal view, pars cephalica strongly elevated in lateral view, anteriorly narrowed to between 0.5 and 0.75 times its maximum width, anterolateral corners with strongly sclerotized, triangular extension, strongly sclerotized lines extend either side of clypeus halfway to elevated portion of pars cephalica; surface of elevated portion of pars cephalica finely reticulate, sides strongly reticulate; nonmarginal pars cephalica setae dark. Clypeus margin unmodified, very strongly curved in front view, low, ALE separated from edge of carapace by less than their radius; setae dark. Eyes: ALE largest, ALE circular, PME circular, PLE oval; posterior eye row straight from above; ALE touching, ALE-PLE touching. Sternum longer than wide, pale orange, uniform; setae dark. Chelicerae, endites, and labium dark red-brown. Chelicerae slightly divergent, anterior face with distal complex projections, largest projection bifurcated (figs. 19A–C, 22C); fang with prominent basal process, broad, fluted outgrowth and large, rounded bump; setae dark. Labium elongated hexagon, not fused to sternum, anterior margin indented at middle, slightly heavier than sternum in sclerotization; with six or more setae on anterior margin. Endites anteromedian tip with one strong, toothlike projection, much more heavily sclerotized than sternum. ABDO- MEN: Ovoid; dorsum soft portions pale white. Book lung covers elliptical. Dorsal scutum brown, covering more than 3/4 of abdomen, more than 1/2 to most of abdomen width, middle surface smooth, sides smooth. Epigastric scutum small lateral sclerites present. Postepigastric scutum pale orange, covering about 1/2 of abdominal length. Dorsum, epigastric area, and postepigastric area setae dark. LEGS: Pale orange, without color pattern; patella plus tibia I shorter than carapace. Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2-0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Palp proximal segments dark red-brown; embolus dark; femur shorter than trochanter, without posteriorly rounded lateral dilation; patella shorter than femur; cymbium dark redbrown; bulb dark red-brown, more than two times as long as cymbium, stout; embolic region obtusely bent, elongate, basal section broad, narrow distally, slightly curved distal tip; fine denticles present on ventral subdistal edge (fig. 21A–C, 22A, B).

FEMALE (PBI_OON 5640, figs. 17D–F, J, 22E; paratype ♀: figs. 20D, F, 21D–G). Total length 1.78. CEPHALOTHORAX: Carapace pars cephalica slightly elevated in lateral view, anteriorly narrowed to 0.49 times its maximum width or less. Clypeus straight in front view. Chelicerae, endites, and labium pale orange. ABDOMEN: Dorsal scutum covering 1/2 to 3/4 of abdomen, between 1/4 and 1/2 abdomen width. Postepigastric scutum widely hexagonal, covering about 1/3 of the abdominal length. LEGS: Leg spination: femora: I p0-1-1; II p0-1-0; tibiae: I, II p2-2- 0; v2-2-0; metatarsi: I, II p1-1-0; v1-1-0. GENITALIA: Ventral view: epigynal region with heavily sclerotized platelike structure, stout, lateral edges straight, epigynal atrium ovoid; convoluted duct thicker than apodemes (figs. 17J, 21D).

OTHER MATERIAL EXAMINED: AUS- TRALIA: Queensland: Mount Lewis Forest Reserve, Mount Lewis Road , 21–22 km from highway, rainforest, leaf litter, 16.54250 ° S, 145.28388 ° E, Apr. 2, 2009 (K. Edward and J. Waldock, QM S95925 View Materials ), 2 ³ GoogleMaps   ; Mount Lewis Forest Reserve, Mount Lewis Road , 21– 22 km from highway, rainforest, leaf litter, 16.54250 ° S, 145.28388 ° E, Apr. 2, 2009 (K. Edward and J. Waldock, QM S95926 View Materials , PBI _ OON 5559), 1 ³ GoogleMaps   ; Black Mountain , 17 km ESE Julatten, rainforest, moss forest litter, 1000 m, 16.65000 ° S, 145.48330 ° E, Apr. 29, 1982 (G. Monteith, D. Yeates, D. Cook, QM S12961 View Materials , PBI _ OON 25809 View Materials ), 1 ♀ GoogleMaps   ; Carbine Tableland, Pauls Luck–Doolins Creek , 1100 m, 16.43333 ° S, 145.26666 ° E, Nov. 30, 1990 (G. Monteith, G. Thompson, D. Cook, H. Janetzki, QM S49371 View Materials , PBI _ OON 22180 View Materials ), 1 ³ GoogleMaps   ; Carbine Uplands area, Mount Lewis Road , rainforest, leaf litter, 820 m, 16.58638 ° S, 145.29250 ° E, Apr. 1, 2009 (K. Staunton, QM S95927 View Materials , PBI _ OON 5576), 1 ♀ GoogleMaps   ; Carbine Uplands area, Mount Lewis Road , rainforest, leaf litter, 600 m, 16.57722 ° S, 145.30916 ° E, Apr. 1, 2009 (K. Staunton, QM S95928 View Materials , PBI _ OON 5580), 1 ³ GoogleMaps   ; Julatten, Mount Lewis Road , Nov. 12, 1975 (A. Walford-Huggins, ANIC, PBI _ OON 5863), 1 ³, 1 ♀   ; Mossman Gorge National Park , circuit track, rainforest, leaf litter, 16.47222 ° S, 145.33055 ° E, May 21, 2007 (K. Edward and K. Pitz, QM S95929 View Materials , PBI _ OON 25723 View Materials ), 1 ³ GoogleMaps   ; Mount Demi north Peak , rainforest, 1050 m, 16.50000 ° S, 145.31670 ° E, Dec. 17, 1995, to Jan. 25, 1996 (G. Monteith, G. Thompson, Ford, QM S43388 View Materials , PBI _ OON 21798 View Materials ), 3 ³, 2 ♀ GoogleMaps   ; Mount Demi summit, 7 km SW Mossman, rainforest, 1100 m, 16.50000 ° S, 145.31670 ° E, Dec. 17, 1995, to Jan. 25, 1996 (G. Monteith, G. Thompson, Ford, QM S44688 View Materials , PBI _ OON 21968 View Materials ), 4 ³, 3 ♀ GoogleMaps   ; Mount Demi , 7 km SW Mossman, rainforest, leaf litter, 1100 m, 16.50000 ° S, 145.31670 ° E, Oct. 29, 1983 (D. Yeates, G. Thompson, QM S16807 View Materials , PBI _ OON 21531 View Materials ), 1 ♀ GoogleMaps   ; Mount Demi , 7 km SW Mossman, rainforest, leaf litter, 950 m, 16.50000 ° S, 145.31670 ° E, Apr. 26, 1983 (G. Monteith, D. Yeates, QM S16074 View Materials , PBI _ OON 25810 View Materials ), 1 ³ GoogleMaps   ; Mount Demi , 7 km SW Mossman, rainforest, leaf litter, 950 m, 16.50000 ° S, 145.31670 ° E, Apr. 26, 1983 (G. Monteith, D. Yeates, QM S16122 View Materials , PBI _ OON 25812 View Materials ), 3 ♀ GoogleMaps   ; Mount Demi , 7 km SW Mossman, rainforest, leaf litter, 1100 m, 16.50000 ° S, 145.31670 ° E, Oct. 29, 1983 (D. Yeates, G. Thompson, QM S16801 View Materials , PBI _ OON 25817 View Materials ), 3 ³, 1 ♀ GoogleMaps   ; Mount Lewis , rainforest, leaf litter, 914 m, 16.58333 ° S, 145.28330 ° E, Sept. 4, 1983 (A. Williamson, QM S12989 View Materials , PBI _ OON 25813 View Materials ), 1 ♀ GoogleMaps   ; Mount Lewis Road, Windmill Creek , rainforest, 900 m, 16.56983 ° S, 145.26750 ° E, Nov. 18, 1997, to Feb. 9, 1998 (G. Monteith, D. Cook, QM S69621 View Materials , PBI _ OON 21965 View Materials ), 1 ♀ GoogleMaps   ; Mount Lewis , 2 km N, rainforest, leaf litter, 1000 m, 16.56667 ° S, 145.26670 ° E, Sept. 9, 1981 (G. Monteith, D. Cook, QM S12962 View Materials , PBI _ OON 25808 View Materials ), 2 ♀ GoogleMaps   ; Mount Spurgeon ( Star house), rainforest, 1100 m, 16.45500 ° S, 145.20660 ° E, Nov. 19, 1997 (D. Clyne, QM S35951 View Materials , PBI _ OON 21769 View Materials ), 1 ♀ GoogleMaps   ; Mount Spurgeon ( Star house), rainforest, 1100 m, 16.45500 ° S, 145.20660 ° E, Nov. 19, 1997 (G. Monteith, QM S43135 View Materials , PBI _ OON 21776 View Materials ), 1 ³ GoogleMaps   ; Mount Spurgeon ( Star house), rainforest, leaf litter, 1100 m, 16.45500 ° S, 145.20660 ° E, Nov. 20, 1997 (G. Monteith, QM S43045 View Materials , PBI _ OON 21790 View Materials ), 1 ♀ GoogleMaps   ; Mount Spurgeon ( Star house), rainforest, leaf litter, 1100 m, 16.45500 ° S, 145.20660 ° E, Nov. 19, 1997 (G. Monteith, QM S43133 View Materials , PBI _ OON 21985 View Materials ), 2 ♀ GoogleMaps   ; Mount Spurgeon, 2 km SE, via Mount Carbine , rainforest, leaf litter, 1100 m, 16.45000 ° S, 145.20000 ° E, Dec. 20, 1988 (G. Monteith, G. Thompson, QM S58138 View Materials , PBI _ OON 22137 View Materials ), 1 ³, 3 ♀ GoogleMaps   ; the Bluff , 11 km W Mossman, rainforest, leaf litter, 950 m, 16.45000 ° S, 145.26670 ° E, Apr. 27, 1983 (G. Monteith, D. Yeates, QM S16072 View Materials , PBI _ OON 25807 View Materials ), 1 ³ GoogleMaps   ; the Bluff , 11 km W Mossman, rainforest, leaf litter, 1000 m, 16.45000 ° S, 145.26670 ° E, Nov. 2, 1983 (G. Monteith, D. Yeates, G. Thompson, QM S16812 View Materials , PBI _ OON 25816 View Materials ), 1 ♀ GoogleMaps   .

DISTRIBUTION: This species is only known from three northern subregions of the Wet Tropics Bioregion (CU, ML, and BM), in northeastern Queensland (map 3).

QM

Queensland Museum

ANIC

Australian National Insect Collection