Coniceromyia crassivena, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 9-10

Coniceromyia crassivena sp. nov.

( Figs. 2E , 8K , 13K , 18 H–I, 24L)

Diagnosis (male). Large wing membrane area between the veins M 2 and CuA 1 (CuA 1 -M 2 /M 2 -M 1 ratio 5.2), M 2 considerably thick ( Fig. 13K ), foretibia with three dorsal setae ( Fig. 2E ).

Material examined. Holotype ♂, BOLIVIA: La Paz: 40km N Caranavi: Cumbre Alto Beni, 15.83ºS, 67.56ºW, 7–14.iv.2004, Malaise trap, 1600m, B. Brown, E. Zumbado col. [LACM ENT 194075].

Description. Male. Body length, 2.8 mm. Head. Frons dark brown, shining, sparsely setulose, with well-defined median furrow. Flagellomere 1 light brown, darker apically, conical-elongate. Arista apical, slightly shorter than flagellomere 1. Palpus yellowish- brown. Gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown; anepisternum bare. Legs yellowish-brown, except midcoxa brown and dorsal macula on posterior face of hind femur. Foretibia with three dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with anterior bare excavation, anteroapical setulose process, group of setae at excavation base including a long curved seta, row of dense setulae on dorsal margin of excavation, and strong seta at base of anteroapical process ( Fig. 8K ). Foremetatarsus ratio 0.28. Midtibia with two anterodorsal and one posterodorsal setae in basal half. Posterior face of hind femur with band of basoventral tiny blunt setulae tapering toward ventral margin ( Figs. 18 H–I). Hind femur height/ length ratio 0.31. Hind tibia with two dorsal setae in basal half. Wing. Darkening in the wing membrane between M 2 and CuA 1 and close to the wing apex; Costa swollen, 0.36 of the wing length; R 2+3 vestigial; M 1 slightly concave; M 2 twice the thickness of M 1, slightly sinuous, CuA 1 -M 2 /M 2 -M 1 ratio 5.2; CuA 1 concave. Halter base brown, apex white. Abdomen. Tergites dark brown, with shining pubescence. Hypopygium light brown ( Figs. 24L ). Left epandrial process bifurcated into a dorsal and a ventral lobe. Ventral lobe with many strong setulae at medial face. Dorsal lobe with subepandrial setulose elevation. Right subepandrial plate expanded posteriorly. Margin of right lobe of hypandrium with lateral projection. Hypoproct with three setae.

Distribution. Known from a single mid-elevation site in Bolivia.

Etymology. Name derived from the Latin roots crassus (thick, stout) and vena, referring to the wing vein M 2 considerably thick.

Figures

FIGURE 2. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. camptoneura, B. C. chrysopa, C. C. circulata, D. C. cladopyga, E. C. crassivena, F. C. dasypoda.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 13. Wings of the males of Coniceromyia new species. A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda.

FIGURE 18. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. camptoneura, C–D. C. chrysopa, E. C. circulata, F–G. C. cladopyga, H–I. C. crassivena, J–K. C. dasypoda.

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.