Coniceromyia diffusa, Ament & Kung & Brown, 2020

Ament, Danilo César, Kung, Giar-Ann & Brown, Brian V., 2020, Forty-one new species of Coniceromyia Borgmeier (Diptera: Phoridae), an identification key, and new distributional records for the species of the genus, Zootaxa 4830 (1), pp. 1-61 : 20-21

Coniceromyia diffusa sp. nov.

( Figs. 3C , 8O , 14C , 19 D–E, 24P, 27H)

Diagnosis (male). Anepisternum bare, left epandrial process with only one discernible process: a subepandrial wall extending from the hypoproct to its apex ( Fig. 24P ), wing without macula ( Fig. 14C ).

Material examined. Holotype ♂, COLOMBIA: Nariño: RN La Planada: Parcela Perm., 1.25°N, 78.25°W, 16.xii–2.i.2001, Malaise trap, 1850m, G. Oliva col. ( IAVH) [LACM ENT 213101] . Paratypes: BOLIVIA: 1♂, La Paz: 40km N Caranavi: Cumbre Alto Beni, 15.83°S, 67.56°W, 14–21.iv.2004, Malaise trap, 1600m, B. Brown, E. Zumbado col. ; 1♂, same but 7–15.iv.2004, S. Gaimari, M. Hauser col.; 2♂, same but 15–19.iv.2003, Brown, Marcotte, Zumbado col. BRAZIL: 1♂, Pará: Carajás: FLONA Carajás , 06°24’08”S 50°20’45”W, 20–27.iv.2010, Malaise trap, Kumagai & Lima col. ( UFMG) . COLOMBIA: 1♂, Valle del Cauca: PNN Farallones de Cali: Alto Anchicaya , 8.43°N, 76.80°W, 10–25.xi.2000, Malaise trap, 900m, S. Sarria col. ; 1♂, Huila: PNN Cueva de Los Guácharos: Alto el Mirador , 1.63°N, 76.10°W, 6–20.i.2002, Malaise trap, 1980m, J. Urbano col. COSTA RICA: 1♂, Puntarenas: Monteverde Biol. Stn.: San Luis, 10.75ºN, 84.75ºW, viii.1994, Malaise trap, Z. Fuentes col. ; 1♂, same but 10.32°N, 84.8°W, 26–28.v.1998, Malaise trap, 1500m, B. Brown & V. Berezovskiy col.; 1♂, Alajuela: Res. Biol. San Ramon , 10.22ºN, 84.62ºW, iv–v.1995, Malaise trap, 900m, P. Hanson col. ; 1♂, same but 23.xi.1994 – 27.i.1995, Malaise trap, 800m, G. Caballo col.; 1♂, Alajuela: Los Chiles , 10.4ºN, 84.33ºW, iii.1990, Malaise trap, 300m, P. Hanson col. ; 1♂, Alajuela: 5km W San Ramon , 10.06ºN, 84.5ºW, iv.1997, Malaise trap, 1200m, O. Castro col. ; 1♂, Guanacaste: Estación Pitilla , 11ºN, 85.43ºW, v.1989, Malaise trap, 600m, P. Hanson col. ; 1♂, Limón: 7km SW Bribri , 9.58ºN, 82.88ºW, ix–x.1989, Malaise trap, P. Hanson col. ; 1♂, San Jose: ZP El Rodeo , 9.91ºN, 84.28ºW, 5–8.viii.2001, Malaise trap, 1000m, Brown, Berezovskiy & Kung col. ; 1♂, Cartago: 4km NE Cañon Genesis II, 9.71°N, 83.91°W, iv.1996, Malaise trap, 2350m, P. Hanson col. ; 1♂, San Isidro de las Peñas Blancas: Texas AT, 20.iv–26.v.2010, Malaise trap, 400m, W. Porras col. ( MZUSP) . ECUADOR: 2♂, Napo: Baeza , 0.6°S, 77.87°W, 15–19.v.1987, Malaise trap, 1550m, B. Brown, L. Coote col. ; 1♂, Pichincha: 17km E Sto Domingo: Tinalandia , 11–13.v.1987, Malaise trap, 800m, B. Brown, L. Coote col. PERU: 3♂, Cusco: 19rd km Quincemil: Rio Araza Tributary , 13°20’S 70°50’W, 23–31.viii.2012, Malaise trap, 874m, R.R. Cavichioli, A. Rafael, A.P.M. Santos & D.M. Takiya col. ( MZUSP) . VENEZUELA: 3♂, Lara: Yacambú , 7.v.1981, 1200m, H.K. Townes col. ; 1♂, Mérida: 3km N La Azulita , 8.72°N, 71.44°W, 8.iv.1988, A. Finnamore & C. Baxfield col.

Description. Male. Body length, 2.3 mm. Head. Frons dark brown, sparsely setulose, with well-defined median furrow. Flagellomere 1 brown, conical-elongate.Arista apical, with length subequal to flagellomere 1 in length. Palpus light brown. Gena with two setae. Thorax. Scutum, scutellum, and pleural regions brown; anepisternum bare. Legs yellowish-brown, foretarsomere 5 brown. Foretibia with two dorsal setae and anterodorsal row of strong setulae. Foremetatarsus with anterior excavation, anteroapical setulose process, group of setae at base of excavation, and seta at base of anteroapical process ( Fig. 8O ). Foremetatarsus ratio 0.35. Posterior face of hind femur with narrow basoventral band of tiny blunt setulae ( Fig. 19 D–E). Hind femur height/length ratio 0.34. Hind tibia with one dorsal seta in basal half. Wing ( Fig. 14C ). Costa swollen, 0.4 of the wing length; R 2 + 3 vestigial; M 1 concave, curved posteriorly at apex; M 2 slightly sinuous, CuA 1 -M 2 /M 2 -M 1 ratio 1.2; CuA 1 sinuous. Halter base brown, apex yellowish-brown to white. Abdomen. Tergites brown, with shining pubescence. Hypopygium light brown ( Fig. 24P ). Left epandrial process with only one discernible process: a subepandrial wall extending from the hypoproct to its apex. Subepandrial wall with setulae at its edge. Margin of right lobe of hypandrium without lateral projection. Hypoproct with two setae.

Distribution. From Costa Rica to Bolivia, occurring in some mid-elevation sites in the Andean region, and in Brazilian Amazonia ( Fig. 27H ).

Etymology. Name derived from the Latin root diffusus (spread out, dispersed), referring to the wide geographical distribution of this species.

Variations. Specimens with a conical flagellomere 1; foretibia with two to three dorsal setae; wing vein R 2+3 present or absent.

Figures

FIGURE 3. Habitus of the holotypes of new species of Coniceromyia, continuation (scale bars=1mm). A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna.

FIGURE 8. Male foremetatarsus of new species of Coniceromyia (scale bars=0.1mm). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga, K. C. crassivena, L. C. dasypoda, M. C. deltopoda, N. C. diadela, O. C. diffusa, P. C. dolichopoda, Q. C. hadrochaeta, R. C. hirtipenna, S. C. inflata, T. C. leia, U. C. litopoda, V. C. luna, W. C. maculata, X. C. megalosoma, Y. C. memorialis, Z. C. niemeyeri, A’. C. numerosa, B’. C. ochyrosticha, C’. C. parallela, D’. C. pectinella.

FIGURE 14. Wings of the males of Coniceromyia new species, continuation. A. C. deltopoda, B. C. diadela, C. C. diffusa, D. C. dolichopoda, E. C. hadrochaeta, F. C. hirtipenna, G. C. inflata, H. C. leia, I. C. litopoda, J. C. luna, K. C. maculata, L. C. megalosoma.

FIGURE 19. Hind femora of the males of Coniceromyia new species, continuation (scale bars on figures of the entire femur=0.3 mm, on figures of ventrobasal part of the femur=0.1 mm). A–B. C. deltopoda, C. C. diadela, D–E. C. diffusa, F–G. C. dolichopoda, H–I. C. hadrochaeta, J. C. hirtipenna, K. C. inflata

FIGURE 24. Hypopygia of Coniceromyia new species in right lateral view (except figure J which is left lateral view). A. C. amazonensis, B. C. anisocrossa, C. C. balbiae, D. C. bellatula, E. C. bisetosa, F. C. brachypoda, G. C. camptoneura, H. C. chrysopa, I. C. circulata, J. C. cladopyga in left lateral view, K. C. cladopyga in right lateral view, L. C. crassivena, M. C. dasypoda, N. C. deltopoda, O. C. diadela, P. C. diffusa, Q. C. dolichopoda, R. C. hadrochaeta, S. C. hirtipenna, T. C. inflata, U. C. leia, V. C. litopoda, W. C. luna, X. C. maculata.

FIGURE 27. Distributional records of Coniceromyia new species (species known from a single or close localities were not illustrated. Squares=holotype, circles=other specimens). A. C. amazonensis, B. C. bellatula, C. C. bisetosa, D. C. brachypoda, E. C. cladopyga, F. C. dasypoda, G. C. deltopoda, H. C. diffusa, I. C. hadrochaeta, J. C. hirtipenna, K. C. litopoda, L. C. luna, M. C. maculata.