Leviana dimidiata (L. Koch, 1871) Framenau & Kuntner, 2022

Framenau, Volker W. & Kuntner, Matjaz, 2022, The new Australian leaf-curling orb-weaving spider genus Leviana (Araneae, Araneidae), Evolutionary Systematics 6 (2), pp. 103-133 : 103

publication ID

https://dx.doi.org/10.3897/evolsyst.6.83573

publication LSID

lsid:zoobank.org:pub:DFF7CEB3-2038-4CA7-B057-CD5BFACC9510

persistent identifier

https://treatment.plazi.org/id/E5F8C8D9-80FF-577A-9F31-E8E8AEDFF071

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scientific name

Leviana dimidiata (L. Koch, 1871)
status

comb. nov.

Leviana dimidiata (L. Koch, 1871) comb. nov.

Figs 1A-D View Figure 1 , 4A, B View Figure 4 , 5A-C View Figure 5 , 6A View Figure 6 , 7A-D View Figure 7 , 8A-F View Figure 8 , 9 View Figure 9

Epeira dimidiata Koch 1871: 95-96, plate 8, figs 1, 1a.

Epeira sylvicola Rainbow 1897: 518-519, plate 17, figs 4, 4a. New synonymy.

Araneus sylvicola (Rainbow).- Rainbow 1909: 222.

Araneus dimidiatus (L. Koch).- Rainbow 1911: 184; Rainbow 1916: 100; Bonnet 1955: 498; Rack 1961: 22.

Araneus sylvicolus (Rainbow).- Rainbow 1911: 194.

Aranea dimidiata (L. Koch).- Roewer 1942: 827.

Aranea sylvicola (Rainbow).- Roewer 1942: 834.

Araneus silvicola (Rainbow).- Bonnet 1955: 598 (corrected spelling; ' silva ' - Latin: wood).

Type material.

Syntypes of Epeira dimidiata L. Koch, 1871: 1 female, Mackay (21°08'S, 149°11'E, Queensland, Australia) (ZMH, ( Rack 1961)-catalogue no. 235) (examined); 2 females, same locality (L. Koch collection) (NHMUK 1915.3.5.594-595) (examined).

Syntypes of Epeira sylvicola Rainbow, 1897: 2 females, Guildford (33°50'S, 150°59'E, New South Wales, Australia) (AM KS.6679) (examined).

Other material examined.

Australia: New South Wales: 2 males, Beecroft, 33°45'S, 151°04'E (AM KS.53874, KS70922); 2 males, 1 female, Beecroft Reserve , 33°45'S, 151°04'E (AM KS.54073, KS.58718, KS.66205) GoogleMaps ; 1 female, Bellevue Hill, Sydney, 33°53'S, 151°17'E (AM KS.70044); 1 male, Bonny Hills Road, off Pacific Highway, S of Port Macquarie , 31°33'S, 152°49'E (AM KS.62864) GoogleMaps ; 1 female, Cheltenham, 33°45'S, 151°05'E (AM KS.56988); 1 female, Como, 34°01'S, 151°04'E (AM KS.32822); 1 female, Cudgen, 28°16'S, 153°33'E (QM); 12 females, 4 juv., Enfield, 33°53'S, 151°06'E (AM KS.33984); 1 male, Forster, 6 km S, 32°12'S, 152°31'E (AM KS.10204); 1 female, Frazer Reserve , Wahroonga, 33°43'S, 151°08'E (AM KS.56766) GoogleMaps ; 1 female, Galston, Crosslands Road, 33°38'S, 151°05'E (AM KS.83701); 1 female, Gosford, 33°25'S, 151°20'E (MV K-10331); 1 female, Jamberoo Mountain , 34°43'S, 151°18'E (AM KS.81993) GoogleMaps ; 1 female, Lake Munmorah State Recreation Reserve , Geebung, 22°12'S, 151°34'E (AM KS.17794) GoogleMaps ; 1 female, same locality (AM KS.17811) GoogleMaps ; 1 male, Maroota State Forest , 33°31'S, 150°59'E (AM KS.73416) GoogleMaps ; 1 female, Medlow Bath, 33°41'S, 150°17'E (AM KS.34097); 1 female, Minnamurra Falls , 34°38'S, 150°44'E (AM KS.100355) GoogleMaps ; 1 female, Moruya, 35°55'S, 150°05'E (AM KS.33981); 2 females, Narara, 33°23'S, 151°21'E (MV K-10332); 1 male, O'Hares Creek , 34°14'S, 150°52'E (AM KS.86691) GoogleMaps ; 1 female, Port Stephens , south shore, 32°42'S, 152°06'E (AM KS.33983) GoogleMaps ; 1 female, Wahroonga, 33°43'S, 151°08'E (AM KS.46010); 1 female, Wallumatta Nature Reserve , North Ryde, 33°48'S, 151°07'E (AM KS.81958) GoogleMaps ; 1 male, Warrumbungles National Park , White Gum carpark area, 31°17'20"S, 149°02'26"E (AM KS.75194) GoogleMaps ; 2 males, 1 female, Warrumbungles National Park , 0.9 km E of western entrance to park, 31°16'35"S, 148°57'49"E (AM KS.75238) GoogleMaps ; 1 female, same locality (AM KS.75239). Queensland GoogleMaps : 1 male, Belmont Hills Bushlands , 27°31'S, 153°07'E (QM S54745 View Materials ) GoogleMaps ; 2 females, same locality (QM S54746 View Materials ) GoogleMaps ; 1 female, same locality (QM S54744 View Materials ) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 3 females, Blackdown Tableland , Stony Creek Falls, 16°53'S, 145°39'E (QM S39784 View Materials ) GoogleMaps ; 1 female, Boondall Wetlands , 27°20'21"S, 153°04'27"E (QM S67469 View Materials ) GoogleMaps ; 3 females, same locality (QM) GoogleMaps ; 3 females, 1 juv., same locality (QM S67470 View Materials ) GoogleMaps ; 1 female, same locality (QM S67468 View Materials ) GoogleMaps ; 1 female, Boonoo Boonoo , 28°53'S, 152°06'E (QM S39797 View Materials ) GoogleMaps ; 1 male, Bribie Island , 27°03'30"S, 153°11'32"E (AM KS.69544) GoogleMaps ; 1 male, same locality (AM KS.69516) GoogleMaps ; 2 males, same locality (AM KS.69609) GoogleMaps ; 2 females, 11 juv., Brisbane , 27°28'S, 153°01'E (QM S39789 View Materials ) GoogleMaps ; 1 female, same locality (QM S69647 View Materials ) GoogleMaps ; 1 female, Buhot Creek , Burbank, 27°35'27"S, 153°10'19"E (QM S67807 View Materials ) GoogleMaps ; 1 male, same locality (QM S67283 View Materials ) GoogleMaps ; 3 males, same locality (QM S67282 View Materials ) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 2 males, 1 female, same locality (QM S67628 View Materials ) GoogleMaps ; 1 males, same locality (QM S111921) GoogleMaps ; 1 male, 3 females, Bulimba Creek , Carindale, 27°30'9"S, 153°06'03"E (QM S67467 View Materials ) GoogleMaps ; 1 female, same locality (QM S67466 View Materials ) GoogleMaps ; 1 female, same locality (QM S67463 View Materials ) GoogleMaps ; 1 male, same locality (QM S67461 View Materials ) GoogleMaps ; 1 female, same locality (QM S67464 View Materials ) GoogleMaps ; 6 males, 6 females, same locality (QM S67462 View Materials ) GoogleMaps ; 1 female, Bundaberg Forest , 24°52'S, 152°21'E (QM) GoogleMaps ; 1 female, Bundaberg , Baldwins Swamp, 24°52'S, 152°21'E (QM S25246 View Materials ) GoogleMaps ; 1 female, Bundaberg, land, 24°52'S, 152°21'E (MV K-10345); 12 females, Camp Milo , Cooloola, 26°00'S, 153°05'E (QM S39750 View Materials ) GoogleMaps ; 1 female, Chelsea Road Bushland Reserve , 27°28'58"S, 153°11'15"E (QM) GoogleMaps ; 2 females, same locality (QM) GoogleMaps ; 4 females, same locality (QM S65837 View Materials ) GoogleMaps ; 2 females, same locality (QM S65840 View Materials ) GoogleMaps ; 1 female, Closeburn , 27°20'S, 152°52'E (QM S39766 View Materials ) GoogleMaps ; 3 females, Consuelo Tablelands, Foleys Yards, 25°01'S, 147°57'E (QM); 2 females, Cooloola , 26°12'S, 153°03'E (QM S39753 View Materials ) GoogleMaps ; 1 female, Cooloola State Forest , 26°12'S, 153°03'E (QM S33393 View Materials ) GoogleMaps ; 1 female, Coombabah Creek 200 m upstream Lakeside Caravan Park, 27°52'S, 153°22'E (QM S39769 View Materials ) GoogleMaps ; 2 females, Curtis Island , 23°37'S, 151°9'E (QM S69916 View Materials ) GoogleMaps ; 1 female, Deepwater National Park , 24°32'S, 151°58'E (QM S25402 View Materials ) GoogleMaps ; 1 female, Downfall Creek , 27°23'S, 153°01'E (QM) GoogleMaps ; 1 female, Drewvale, Illaweena Street, 27°38'39"S, 153°03'47"E (QM); 1 female, same locality (QM) GoogleMaps ; 5 females, same locality (QM S65839 View Materials ) GoogleMaps ; 1 female, Eidsvold , 25°22'S, 151°07'E (QM S39765 View Materials ) GoogleMaps ; 1 female, Enoggera Army land, Brisbane, 27°26'S, 152°59'E (QM S22429 View Materials ) GoogleMaps ; 1 female, Expedition Range National Park , 25°12'S, 148°59'E (QM S44180 View Materials ) GoogleMaps ; 1 male, 5 females, 9 juv., Finch Hatton National Park , 21°9'S, 148°38'E (QM) GoogleMaps ; 1 female, Forty Mile Scrub, SW Mt Garnet , 18°05'S, 144°51'E (QM S39778 View Materials ) GoogleMaps ; 1 female, Girraween National Park , near, 28°50'S, 151°55'E (QM S39774 View Materials ) GoogleMaps ; 1 female, Gold Creek Reservoir, 27°27'53"S, 152°52'32"E (QM S54749 View Materials ) GoogleMaps ; 1 female, same locality (QM S54750 View Materials ) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 1 female, same locality (QM S54751 View Materials ) GoogleMaps ; 1 female, same locality (QM S54748 View Materials ) GoogleMaps ; 3 females, same locality (QM S54747 View Materials ) GoogleMaps ; 1 female, Hann Tableland , northern end, 16°48'S, 145°10'E (QM S40510 View Materials ) GoogleMaps ; 1 female, same locality (QM S41332 View Materials ) GoogleMaps ; 3 females, same locality (QM S40498 View Materials ) GoogleMaps ; 2 females, Helidon , 27°33'S, 152°08'E (QM S39795 View Materials ) GoogleMaps ; 1 female, Hurdie Gully , 14.8 km WSW Monto, 24°55'S, 150°59'E (QM S44168 View Materials ) GoogleMaps ; 1 female, Ipswich , 27°37'S, 152°47'E (QM S39763 View Materials ) GoogleMaps ; 2 females, Karawatha Forest , 27°37'33"S, 153°05'24"E (QM S67460 View Materials ) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 2 males, same locality (QM S67459 View Materials ) GoogleMaps ; 1 female, Kenniffs Cave , Mt Moffat, 25°01'S, 147°57'E (QM) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 1 female, Kroombit Tops, 24°22'S, 151°01'E (QM); 1 female, same locality (QM) GoogleMaps ; 4 females, same locality (QM S39751 View Materials ) GoogleMaps ; 2 females, Kroombit Tops, Lower Dry Creek , 24°24'S, 151°01'E (QM) GoogleMaps ; 7 females, Kroombit Tops, Upper Dry Creek , 24°25'S, 151°03'E (QM) GoogleMaps ; 4 females, Kroombit Tops, Northern Escarpment , 24°22'S, 151°01'E (QM) GoogleMaps ; 1 female, Kroombit Tops, Upper Kroombit Creek , 24°25'S, 151°03'E (QM) GoogleMaps ; 1 female, Kroombit Tops, Upper TA47 Creek, 24°25'S, 151°03'E (QM); 1 female, Kuraby , 27°37'S, 153°05'E (QM S41023 View Materials ) GoogleMaps ; 1 male, 1 female, Mahogany Forest , Mt Moffat National Park, 24°56'S, 148°04'E (QM S111925) GoogleMaps ; 2 females, Molangdool, 24°45'S, 151°33'E (AM KS.33985); 1 female, Moreton Island , 27°11'S, 153°24'E (QM S39782 View Materials ) GoogleMaps ; 15 females, Mt Abbott , 20°06'S, 147°45'E (QM S38572 View Materials ) GoogleMaps ; 1 female, Mt Coolum , 26°34'S, 153°05'E (QM S111923) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 1 female, Mt Flinders , 22°32'S, 150°46'E (QM S39768 View Materials ) GoogleMaps ; 1 female, Mt Garnet , 17°41'S, 145°07'E (QM) GoogleMaps ; 1 female, Mt Moffat National Park , 25°01'S, 147°57'E (QM) GoogleMaps ; 1 female, Mt Moffat , summit, 25°01'S, 147°57'E (QM) GoogleMaps ; 1 female, 1 juv., Mt Moffat , top camp, 25°01'S, 147°57'E (QM S111924) GoogleMaps ; 2 males, 2 females, 3 juv., Nelson, 12°38'S, 143°15'E (AM KS.32825); 7 females, Nerimberah, via Rockhampton, 23°24'S, 150°35'E (QM S9923); 1 female, North Stradbroke Island , 27°35'S, 153°27'E (QM S55768 View Materials ) GoogleMaps ; 2 females, same locality (QM S55769 View Materials ) GoogleMaps ; 3 females, same locality (QM S55744 View Materials ) GoogleMaps ; 7 females, same locality (QM S55632 View Materials ) GoogleMaps ; 2 females, same locality (QM S55743 View Materials ) GoogleMaps ; 4 females, same locality (QM S56284 View Materials ) GoogleMaps ; 2 females, same locality (QM S55631 View Materials ) GoogleMaps ; 1 female, same locality (QM S55745 View Materials ) GoogleMaps ; 1 female, same locality (MV K-10333) GoogleMaps ; 1 female, Orchid Beach , Fraser Island, 24°58'S, 153°19'E (QM S31029 View Materials ) GoogleMaps ; 2 females, Peel Island , 27°30'S, 153°21'E (QM S39748 View Materials ) GoogleMaps ; 1 female, Percy Island , 21°24'S, 150°20'E (QM S39787 View Materials ) GoogleMaps ; 2 females, Ransome Reserve , 27°29'34"S, 153°11'05"E (QM) GoogleMaps ; 1 male, same locality (QM) GoogleMaps ; 4 females, same locality (QM) GoogleMaps ; 2 females, same locality (QM S65838 View Materials ) GoogleMaps ; 3 males, 1 female, same locality (QM S65841 View Materials ) GoogleMaps ; 1 female, Rochedale , 27°37'S, 153°9'E (QM S39757 View Materials ) GoogleMaps ; 1 female, same locality (QM S39760 View Materials ) GoogleMaps ; 3 males, 6 females, 2 juv., Rochedale State Forest , 27°37'S, 153°09'E (QM S39754 View Materials ) GoogleMaps ; 1 female, same locality (QM S39759 View Materials ) GoogleMaps ; 1 female, same locality (QM S39758 View Materials ) GoogleMaps ; 1 female, same locality (QM S39755 View Materials ) GoogleMaps ; 1 female, same locality (QM) GoogleMaps ; 2 females, same locality (QM S39756 View Materials ) GoogleMaps ; 1 female, Rundle Shale camp site, 23°39'S, 150°59'E (QM S39781 View Materials ) GoogleMaps ; 1 female, South Percy Island , Central Valley, 21°45'S, 150°19'E (QM S33318 View Materials ) GoogleMaps ; 1 female, South Percy Island , NW Bay, 21°45'S, 150°19'E (QM) GoogleMaps ; 2 females, Stanthorpe , 28°40'S, 151°56'E (QM S39783 View Materials ) GoogleMaps ; 1 male, Taringabah , 17°43'S, 145°56'E (QM S39770 View Materials ) GoogleMaps ; 1 female, Teewah Creek , Cooloola, 26°05'S, 153°02'E (QM S39788 View Materials ) GoogleMaps ; 1 male, 3 females, 1 juv., Teviot Brook , Boonah, 27°55'S, 152°33'E (QM S39762 View Materials ) GoogleMaps ; 1 female, Top Moffat Camp, Mt Moffat National Park , 25°01'S, 147°05'E (QM S111926) GoogleMaps ; 1 female, Upper Brookfield , 27°30'S, 152°55'E (QM S39796 View Materials ) GoogleMaps ; 1 female, Warro , via Lowmead, 26°56'S, 150°55'E (QM S39779 View Materials ) GoogleMaps ; 1 female, Woodgate , 25°06'S, 152°34'E (QM S39780 View Materials ) GoogleMaps ; 10 females, Yeppoon , Emu Park, Rockhampton, 23°16'S, 150°05'E (QM S69646 View Materials ) GoogleMaps ; 1 female, same locality (QM S39761 View Materials ) GoogleMaps ; Victoria: 2 females, Bruthen, 37°42'S, 147°50'E (MV K-10335); 2 females, Dermody's Camp , Avon River State Forest, 37°48'17"S, 146°55'03"E (WAM T68049 View Materials ) GoogleMaps .

Diagnosis.

Leviana dimidiata comb. nov. is most similar to L. folium sp. nov. in both somatic (size, colouration) and genitalic morphology. However, they can be clearly separated by the abdominal pattern, particularly by the shape of the black anterior shoulder marks and lateral lines. In L. dimidiata comb. nov., the anterior black marks and lateral lines form an obtuse internal angle (e.g. Figs 7A View Figure 7 , 8A View Figure 8 - arrows), whereas in L. folium sp. nov. this angle is acute (e.g. Figs 12A View Figure 12 , 13A View Figure 13 - arrows). Male pedipalp morphology differs subtly in some key sclerites; for example, the terminal apophysis has a sclerotised tip in L. dimidiata comb. nov. which is absent in L. folium sp. nov. (Fig. 7D View Figure 7 vs Fig. 13D View Figure 13 ), the conductor is more prominent and more elongated and the spine in the basal arch of the median apophysis is comparatively much larger (Fig. 7C View Figure 7 vs Fig. 12C View Figure 12 ). The base of the scape of the female epigyne protrudes further anteriorly in L. dimidiata comb. nov. than in L. folium sp. nov. (Fig. 8C View Figure 8 vs Fig. 13C View Figure 13 ).

Description.

Male (based on QM S111921). Total length 6.6. Carapace (Fig. 7A View Figure 7 ) 3.4 long, 2.3 wide; light brown centrally with dark brown lateral flanks; white setae mainly in cephalic and centrally lighter area; fovea indistinct roundish depression. Eyes: AME 0.20, ALE 0.13, PME 0.14, PLE 0.14; row of eyes: AME 0.48, PME 0.44, PLE 1.12. Sternum (Fig. 7B View Figure 7 ) 1.6 long, 1.1 wide; orange-brown; heavy dark grey pigmentation towards margins; few white setae; few brown bristles marginally. Labium subtriangular, 0.48 long, 0.51 wide; basally dark reddish-brown. Maxillae yellow-brown. Chelicerae dark-brown; few white setae in basal half; three retromarginal teeth of equal size, four promarginal teeth with the second-basal one largest. Legs: leg formula I> II> IV> III; dark brown with distinct yellow-brown annulations; lengths of segments (femur + patella + tibia + metatarsus + tarsus = total leg length): pedipalp 0.8 + 0.2 + 0.2 + - + 1.0 = 2.2, I 3.5 + 1.5 + 3.2 + 3.4 + 1.3 = 12.9, II 2.9 + 1.4 + 2.4 + 2.9 + 1.1 = 10.7, III 1.9 + 0.8 + 1.1 + 1.3 + 0.8 = 5.9, IV 2.4 + 1.1 + 2.8 + 2.9 + 0.8 = 8.1. Abdomen (Fig. 7A, B View Figure 7 ) 3.7 long, 2.4 wide; elongate ovoid; dorsally yellow-grey becoming darker posteriorly; two black central marking anteriorly that continue laterally as black lines almost to spinnerets; four pairs of dark brown sigilla centrally; covered with long brown bristles and white setae; venter centrally dark grey with two longitudinal lateral lighter bands, covered with brown setae; spinnerets light brown.

Pedipalps (Figs 7C, D View Figure 7 ; also 4A, B, 5A-C): median apophysis apically broadly lobed, basally with thorn opposing a hooded base; terminal apophysis fleshy, elongated with double-tip; embolus with broad base, curved and short tip; conductor wider than high in ventral view.

Female (based on QM S67628; epigyne variation QM S25402). Total length 7.1. Carapace (Figs 8A View Figure 8 ) 3.5 long, 2.6 wide; light yellowish-brown, cephalic area somewhat darker and with a dark brown median band; few white setae in particular in cephalic area; fovea shallow and poorly demarcated. Eyes: AME 0.18, ALE 0.15, PME 0.13, PLE 0.11; row of eyes: AME 0.46, PME 0.35, PLE 1.23. Sternum (Fig. 8B View Figure 8 ) 1.8 long, 1.3 wide; yellowish-brown; dark brown pigmentation laterally; covered with few white setae; few long brown bristles marginally. Labium 0.64 long, 0.44 wide, colouration as male. Maxillae as male. Chelicerae dark brown; few white setae in basal half, black setae mesally mainly in apical half; four promarginal teeth, with the apical and third one largest; three retromarginal teeth of similar size. Legs: leg formula I> II> IV> III; light brown with indistinct darker annulations, most distinct on tibiae and metatarsi of leg III and IV; lengths of segments (femur + patella + tibia + metatarsus + tarsus = total length of leg): pedipalp 1.0 + 0.3 + 0.2 + - + 0.8 = 2.3, I 2.9 + 1.5 + 2.3 + 2.4 + 1.1 = 10.2, II 2.6 + 1.1 + 1.9 + 2.1 + 0.8 = 8.5, III 1.6 + 1.0 + 1.5 + 1.1 + 0.6 = 5.8, IV 2.7 + 1.1 + 1.8 + 1.9 + 0.8 = 8.3. Abdomen (Fig. 8A, B View Figure 8 ) 3.5 long, 3.2 wide; ovoid, slightly dorsoventrally compressed; dorsal colour pattern and setae largely as male, however lighter; venter as male, but with narrow light guanine line laterally; spinnerets brown.

Epigyne (Fig. 8C-F View Figure 8 ) ovoid wider than long, atrium bulging; central division broadly oval with narrow base; scape with almost parallel lateral border, almost translucent; often broken off anteriorly; spermathecae ovoid, almost touching centrally; fertilization duct very slightly curved and attaching laterally at atrium (Fig. 6A View Figure 6 ).

Variation.

Males total length 4.8-6.6 (n = 5); females total length 6.4-9.9 (n = 7). There is little colour variation in males and females of this species, although the posterior part of the abdomen might be darker than in the specimens illustrated. Two out of seven females measured for this study had their epigyne scape intact.

Remarks.

Rack (1961) considered a female in the ZMH the holotype of Epeira dimidiata , however, two further females from the type locality lodged in the NHMUK clearly bear the typical labels of L. Koch which suggests these to by syntypes. The original description does not state the number of specimens on which the description was based, and we therefore consider all specimens from Port McKay in the ZMH and NHMUK as syntypes.

The syntypes of Araneus sylvicolus (Rainbow, 1897) are clearly conspecific with those of L. dimidiata comb. nov. based on genitalic and somatic morphology, considering the intraspecific variation as described here. We therefore consider E. sylvicola Rainbow, 1897 a junior synonym of L. dimidiata (L. Koch, 1871) comb. nov.

Life history and habitat preferences.

Adult males were mainly found between September and January, with a peak in November. Mature females were mainly found between September and April, with two peaks in December and February.

Records of L. dimidiata comb. nov. are from a variety of open and closed forests and woodlands, including those of eucalypt, Casuarina and Melaleuca . However, there are also some records from heathlands and vine scrubs.

Distribution.

Eastern Australia from far north Queensland to eastern Victoria (Fig. 9 View Figure 9 ).

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Araneidae

Genus

Leviana

Loc

Leviana dimidiata (L. Koch, 1871)

Framenau, Volker W. & Kuntner, Matjaz 2022
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Loc

Epeira dimidiata

Framenau & Kuntner 2022
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Loc

Epeira sylvicola

Framenau & Kuntner 2022
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Loc

Araneus sylvicola

Framenau & Kuntner 2022
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Loc

Araneus dimidiatus

Framenau & Kuntner 2022
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Loc

Araneus sylvicolus

Framenau & Kuntner 2022
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Loc

Aranea dimidiata

Framenau & Kuntner 2022
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Loc

Aranea sylvicola

Framenau & Kuntner 2022
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Araneus silvicola

Framenau & Kuntner 2022
2022