Pison tegulare Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 445-448

Pison tegulare Pulawski , species nova

Figures 1084-1091.

NAME DERIVATION.– Tegulare is a Latin neuter adjective derived from tegula, which is unusually long in this species.

RECOGNITION.– Pison tegulare shares with P. curiosum an unusually long tegula, extending beyond the anterior margin of the axilla. It differs in having the tegula nearly completely punctate and setose (only a narrow, marginal rim is impunctate and asetose), with the inner margin concave posteriorly and the yellowish brown mandible (except basally and apically), and in the female the lower gena and the forefemur with a psammophore and the lower gena impunctate and asetose between the oral fossa and the psammophore. In P. curiosum , the tegula is largely impunctate and asetose, with the inner margin convex posteriorly, the mandible is black except brown apically, and in the female the lower gena and the forefemur have no psammophore, and the lower gena is punctate and setose on each side of the oral fossa.

Two undescribed forms have the tegula identical as in tegulare ; they may be individual variants of tegulare or closely related species. They differ as follows:

a. clypeal lip of female markedly more prominent (1 ♀ from 23 km SE Cobar, New South Wales, ANIC; 1 ♀ from Karijini National Park , Western Australia, ANIC) .

b. male flagellum longer, e.g., dorsal length of flagellomere II is 2.2 × its apical width, rather than 1.9 × (1 ♂, Kakadu National Pak , Northern Territory, ANIC) .

DESCRIPTION.– Frons dull, punctures compressed against each other, middle supraantennal carina largely replaced by flat line. Occipital carina narrowly separated from hypostomal carina. Gena narrow in dorsal view. Labrum not emarginate. Anteromedian pronotal pit transversely elongate, slightly longer than midocellar diameter. Propleuron impunctate or nearly so over most of its surface. Scutum not foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal punctures well defined, averaging less than one diameter apart (several punctures may be more than one diameter apart); interspaces unsculptured (Fig. 1087). Tegula conspicuously elongate, extending beyond anterior margin of axilla, with inner margin concave posteriorly, punctate throughout except for narrow marginal rim (Fig. 1087). Mesopleural punctures compressed against each other, partly concealed by vestiture. Postspiracular carina present, varying from slightly longer than half midocellar diameter to about as long as midocellar diameter. Metapleural sulcus costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum closely punctate (interspaces in many specimens forming minute ridges), with longitudinal ridges basally, and in some specimens with short transverse ridges emerging from middle carina; side punctate, interspaces confluent into small ridges; posterior surface transversely ridged, punctate between ridges. Posteroventral forefemoral surface closely punctate, punctures nearly compressed against each other. Hindcoxal dorsum with outer margin not carinate. Punctures of tergum I, on horizontal part, varying from less than to more than one diameter apart. Punctures of sternum II several diameters apart medially, minuscule in most specimens, but some punctures larger in female from Mount Augustus National Park, Western Australia, and all punctures larger in male from the same Park; apical depression impunctate mesally.

Setae silvery, appressed on frons, scutum, and tergum I, oriented ventrolaterally on frons; largely concealing integument on clypeus (except lamella); lower gena in male with several suberect setae that are about as long as midocellar diameter (see below for female). Apical from all ferruginous to all black.

♀.– Upper interocular distance equal to 0.76 × lower interocular distance; ocellocular distance equal to 0.7-0.8 × hindocellar diameter, distance between hindocelli equal to 1.1-1.4 × hindocellar diameter; eye height equal to 0.98-1.04 × distance between eye notches. Free margin of clypeal lamella arcuate, lamella varying significantly from narrow (Fig. 1085) to wide (Fig. 1084). Dorsal length of flagellomere I 2.1-2.2 × apical width, of flagellomere IX 1.0-1.2 × apical width. Lower gena, mandibular posterior margin, propleural outer margins, and forefemoral venter with psammophores (longest setae of genal, mandibular, and forefemoral psammophores about 0.6-0.7 ×, 0.5-0.8 ×, and 0.7-1.0 ×, respectively, of greatest forefemoral width); lower gena shiny, unsculptured, and asetose between oral fossa and psammophore. Mandible: trimmal carina without incision. Length 5.1-5.3 mm; head width 1.6-1.9 mm.

♂.– Upper interocular distance equal to 0.84 × lower interocular distance; ocellocular distance equal to 1.0-1.7 × hindocellar diameter, distance between hindocelli equal to 1.2-2.0 × hindocellar diameter; eye height equal to 0.96-1.0 × distance between eye notches. Free margin of clypeal lamella acutely angulate (Fig. 1086). Flagellomeres III and IV slightly convex ventrally. Dorsal length of flagellomere I 1.7 × apical width, of flagellomere X 0.8-0.9 × apical width. Sternum VIII gently rounded apically, not emarginate, without posterolateral corner (Fig. 1088). Genitalia: Figs. 1089, 1090. Length 4.5-4.6 mm; head width 1.6 mm.

GEOGRAPHIC DISTRIBUTION (Fig. 1091).– New South Wales, Northern Territory, Queensland, South Australia, Western Australia.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA: Western Australia: Merredin , 12-13 Dec 1935, R.E. Turner ( BMNH).

PARATYPES: AUSTRALIA: New South Wales :

Barnatos Tank 56 mi. W Cobar, 1 Jan 1966, O.W.

Richards (1 ♀, BMNH); 23 km SE Cobar at 31°31ʹS

146°06ʹE, 3 Dec 1981, I.D. Naumann and J.C.

Cardale (1 ♀, ANIC); Coonabarabran, 1 Dec 1991 ,

N.W. Rodd (1 ♀, AMS); 40 km E Gol Gol , 27 Nov

1992, N.W. Rodd (1 ♀, AMS); Springs Creek 68 km

SW Wilcannia at 31°44ʹS 142°41ʹE, 29 Nov 1981,

J.C. Cardale and I.D. Naumann (1 ♀, ANIC) .

Northern Territory: Alice Springs, 5 Oct 1972, Z.

Liepa (1 ♀, ANIC); Burulba Billabong in Kakadu

National Park, 27 June 1980, I.D. Naumann (1 ♀,

1, ♂ ANIC); West MacDonnell National Park ca

3 km W road to Simpson Gap at 23°41.8ʹS

133°41.7ʹE, Ch.M. Palmer, 27 Aug – 27 Sept 2007 FIGURE 1091. Collecting localities of Pison tegulare (1 ♀, CAS), 27 Sept – 27 Oct 2007 (1 ♂, CAS), Pulawski, sp. nov .

27 Oct – 27 Nov 2007 (2 ♂, NTM). Queensland: Amby , 22-27 Nov 1979, H.E. and M.A. Evans and A. Hook (1 ♀, 2 ♂, QMB); 3 km NE Mount Webb at 15°03ʹS 145°09ʹE , 1-3 Oct 1980. J.C. Cardale (1 ♀, ANIC). South Australia: Brachina Gorge in Flinders Ranges National Park at 31°20ʹS 138°34ʹE , 4-10 Nov 1987, I.D. Naumann and J.C. Cardale (1 ♂, CAS); Chowilla Game Reserve 24 air km N Renmark at 34°00.0ʹS 140°49.4ʹE, 2, 3, 5, and 6 Dec 2010, V. Ahrens and W.J. Pulawski (4 ♂, CAS); Dingly Dell Camp on Oraparinna Creek at 31°21ʹS 138°42ʹE , 4-10 Nov 1987, I.D. Naumann and J.C. Cardale (1 ♀, 1 ♂, ANIC); Everard Park Station (now Mimili): creek near Victory Well , 20 Oct 1970, G. Gross and E. Matthews (1 ♀, SAM); Mount Davies and vicinity , 18-21 Oct 1972, H.E. Evans (2 ♂, ANIC; 1 ♂, CAS); Point Sinclair 19 km S Penong at 32°05.0ʹS 132°59.0ʹE , 12 Jan 2011, V. Ahrens and W.J. Pulawski (1 ♀, CAS); Quinyambie Station 5.2 km S Coonanna Bore at 29°53ʹ29ʺS 140°47ʹ21ʺE , 27 Oct – 1 Nov 2008, Waterhouse Survey, no collector (1 ♂, SAM); Quinyambie Station 22 km NE Coonanna Bore at 29°41ʹ58ʺS 140°55ʹ56ʺE , 26-31 Oct 2008, Waterhouse Survey, no collector (3 ♀, 2 ♂, SAM); Quinyambie Station 23.2 km NE Coonanna Bore at 29°42ʹ07ʺS 140°56ʹ07ʺE , Waterhouse Survey, no collector, 26-31 Oct 2008 (1 ♀, CAS) and 27 Oct – 1 Nov 2008 (1 ♀, SAM); 14 km WNW Renmark at 34°07ʹS 140°37ʹE , 13 Dec 1995 – 25 Jan 1996, K.R. Pullen (1 ♀, CAS); 79 km NNW Renmark at 33°31ʹS 140°29ʹE , 24 Jan – 20 Feb 1996, K.R. Pullen (1 ♀, CAS). Western Australia: ca 11 km N Jurien Bay at 30°12.4ʹS 115°00.4ʹE , 1 Nov 2008, V. Ahrens and W.J. Pulawski (1 ♂, CAS); Karijini National Park at 22°26.3ʹS 118°22.9ʹE, M.E. Irwin, and F.D. Parker (1 ♀, CAS); Mount Augustus National Park , M.E. Irwin and F.D. Parker, at 24°18.0ʹS 116°47.6ʹE , 25 Apr – 7 May 2003 (1 ♀, ANIC) and 24°22.8ʹS 116°54.2ʹE , 9-22 May 2003 (1 ♂, CAS); 158 km S Newman (= 9 km N Kumarina Roadhouse ) at 24°37.8ʹS 119°36.8ʹE , 7-18 May 2003, F.D. Parker and M.E. Irwin (1 ♂, ANIC) .