Pison clypeare Pulawski, 2018

Pulawski, Wojciech J., 2018, A Revision of the Wasp Genus Pison Jurine, 1808 of Australia and New Zealand, New Guinea, and the Pacific Islands (Hymenoptera: Crabronidae), Proceedings of the California Academy of Sciences 65, pp. 1-584 : 125-128

Pison clypeare Pulawski , species nova

Figures 241-247.

NAME DERIVATION.– Clypeare is a Neolatin neuter adjective derived from clypeus; with reference to the unusual clypeus of this species.

RECOGNITION.– Pison clypeare has two submarginal cell, the second one elongate, with length of posterior margin equal to 1.9-2.0 × its height ( Fig. 244 ), a partly impunctate tegula, the scutellum foremargin with a foveate sulcus between the axillae (sulcus inconspicuous in some specimens), the propodeal dorsum and posterior surface separated from the side by a well-defined carina, and the propodeal side concave. The female (the male is unknown) resembles P. noctulum in having the clypeal free margin not differentiated into the lobe and lateral sections, forming a single arcuate line orbit to orbit ( Fig. 241 ). Unlike P. noctulum , the punctures of the upper frons average about one diameter apart in P. clypeare , the propodeal dorsum is coarsely rugose posterolaterally, the posterior propodeal surface is coarsely ridged throughout, the wing veins are ferruginous to yellowish, and the length of female flagellomere I is 1.8 × its apical width. In P. noctulum , the punctures of the upper frons are less than one diameter apart, the propodeal dorsum is finely sculptured posterolaterally, the posterior propodeal surface has fine ridges that become evanescent dorsally, the wing veins are dark brown, and the length of female flagellomere I is 1.5 × the apical width.

DESCRIPTION.– Frons swollen above antennal socket, finely punctate, punctures averaging about one diameter apart, interspaces conspicuously microsculptured; middle supraantennal carina absent. Midocellus smaller than hindocellus. Gena narrow in dorsal view ( Fig. 242 ). Labrum shallowly emarginate. Anteromedian pronotal pit transversely elongate, about 1.5 × as long as midocellar diameter. Propleuron impunctate anteriorly. Scutum slightly foveate along flange, without longitudinal ridges adjacent to posterior margin; scutal and mesopleural punctures well defined, less than one diameter apart ( Fig. 243 ), interspaces microsculptured, dull. Tegula enlarged, its outer margin almost rectilinear in anterior half. Postspiracular carina absent. Scutellum foremargin with foveate sulcus between axillae (sulcus inconspicuous in some specimens). Metapleural sulcus conspicuously costulate between dorsal and ventral metapleural pits. Propodeum with irregular longitudinal carina separating side from dorsum and posterior surface and extending from gastral socket area toward spiracle; dorsum with well-defined, irregular, transverse ridges ( Fig. 245 ), oblique near base, ridges anastomosed and with punctures between them, extending laterally to lateral longitudinal carina, becoming coarsely rugose at posterolateral angle ( Fig. 246 ); side concave, ridged, punctate between ridges; posterior surface conspicuously, transversely ridged, ridges connected by fine vertical carinae. Forewing with two submarginal cells ( Fig. 244 ); second submarginal cell broad (length of posterior margin 1.9-2.0 × its height). Posteroventral forefemoral surface with well defined punctures 1-2 diameters apart; interspaces microsculptured. Outer surface of hindtibia with evanescent spines. Punctures of tergum I well defined, less than one diameter apart to about one diameter apart. Sterna punctate throughout.

Setae silvery, appressed on head, mesothorax, forecoxal venter, femoral venters, and tergum I (setae of lower gena subappressed, slightly curved, about as long as 0.7 × midocellar diameter), oriented dorsally on frons, not concealing integument on clypeus. Apical depressions of terga with inconspicuous, silvery, setal fasciae.

Head, thorax, propodeum, and gaster black except mandible brown apically. Wing membrane slightly yellowish and veins light brown. Legs all black in specimen from Coocumbac Island and Karawatha Forest, tarsi ferruginous in specimen from Mackay, and tibiae and tarsi ferruginous in specimens from Coen, Granite Gorge, and Split Rock. Hindtibial spurs brown.

♀.– Upper interocular distance equal to 0.76-0.82 × lower interocular distance; ocellocular distance equal to 0.7-0.8 × hindocellar diameter, distance between hindocelli equal to 1.3 × hindocellar diameter; eye height equal to 1.02-1.06 × distance between eye notches. Middle clypeal lobe not differentiated, free clypeal margin evenly arcuate ( Fig. 241 ) to slightly sinuous. Dorsal length of flagellomere I 1.8 × apical width, of flagellomere IX 0.9 × apical width. Mandible: trimmal carina with small incision shortly beyond midlength. Length 7.2-7.3 mm; head width 1.7 mm.

♂.– Unknown.

GEOGRAPHIC DISTRIBUTION ( Fig. 247 ).– Eastern New South Wales, eastern Queensland.

RECORDS.– HOLOTYPE: ♀, AUSTRALIA:

Queensland: Mackay, Feb 1899, R. Turner

(BMNH).

PARATYPES: AUSTRALIA: New South Wales :

Coocumbac Island near Taree, 21-29 Nov 1994,

G. and T. Williams (1 ♀, ANIC). Queensland: Brisbane: Karawatha Forest at 28°38.6ʹS 153°04.2ʹE ,

12 Dec 2006, W.J. Pulawski (1 ♀, CAS); Coen at

13°57ʹS 143°12ʹE, 16 Aug – 13 Sept 1993, P. Zborowski and S. Shattuck (1 ♀, ANIC); Granite Gorge ca 6 km SW Mareeba, 20 Jan 1999, M. Generani and

P.L. Scaramozzino (1 ♀, CAS); Split Rock at

15°39ʹS 144°31ʹE, 24 June – 29 July 1992, P. Zborowski and E.S. Nielsen (1 ♀, CAS) and

28 May – 26 June 1993, P. Zborowski and I.D. Nau- FIGURE 247 . Collecting localities of Pison clypeare mann (1 ♀, ANIC). Pulawski, sp. nov.

Figures

FIGURES 197–299. Pison bimbi Pulawski, sp. nov., female. (197) Clypeus and mandibles; (198) Head in dorsal view; (199) Pronotum in dorsal view. FIGURE 200. Collecting localities of Pison bimbi Pulawski, sp. nov.