Draposa oakleyi, Kronestedt, Torbjörn, 2010

Kronestedt, Torbjörn, 2010, Draposa, a new wolf spider genus from South and Southeast Asia (Araneae: Lycosidae), Zootaxa 2637, pp. 31-54 : 45-48

Draposa oakleyi (Gravely, 1924) comb. nov.

Figs 3, 4, 10, 11, 21, 22, 25, 31, 44-49

Pardosa oakleyi Gravely , 1924: 610, fig. 5C (♂♀). Tikader & Malhotra 1980: 337-338, figs 179-182 (♂♀); Okuma et al. 1993: 51, fig 45B (♀).

Pardosa laborensis Dyal , 1935: 145-146, pl. 13 figs 43-44 (♂♀). Sadana 1971: 107, fig. 3 0). Synonymy established in Tikader & Malhotra (1980).

Type material. Syntypes of Pardosa oakleyi Gravely , 1924: 1♂, 1♀, India, Tamil Nadu, Ootacamund (ca 11°24'N 76°41'E) ( National Zoological Collection, Kolkata), not examined. 1♀, India, Nilgiris, Ootacamund , 6400-8000 ft, 20-30 May 1921 (Gravely, BM 1924.V.13.20-21 ) ( BMNH, ex Madras Museum), examined. Syntypes of Pardosa lahorensis Dyal , 1935: depository unknown .

Other material examined. INDIA. Chandigarh: Chandigarh (ca 30°44'N 76°46'E), from grassy ground, 14 October 1965 ( BMNH), 2♂ 2♀ . Punjab: Ludhiana (ca 30°54'N 76°51'E), 10 April 1974 (G. L. Sadana. NHRS), 1♀ ; Patiala City (30°21'N 76° 27'E), University Campus , 3-8 May 1999 (Y. M. Marusik, NHRS, ZMUM), 8♂ 6♀ ; d:o, 24-25 June 1999 Y. M. Marusik, ZMUM), 2♂ . Tamil Nadu: Nilgiris, Ootacamund (ca 11°24''N 76°41'E), 2450 m asl, grassy mountain slope, 22 April 1979 (P. T. Lehtinen, NHRS, ZMUT), 2♂ 7♀ . BANGLADESH.: Rajshahi: Rajshahi (ca 24°23'N 88°37'E), grassy dry lawn, 4 November 1973 (A. Islam, ZMUT), 4♀ .

Diagnosis. Male distinguished by configuration of palp, notably shape of tegular apophysis with two close acute protrusions (Fig. 10; cf. D. atropalpis : Fig. 8); female by shape of epigyne, notably long median septum almost covering opening of median cavity (Fig. 11; cf. D. atropalpis : Fig. 9).

Description. Male (from Ootacamund, India). Total length 4.4, carapace 2.55 long, 1.90 wide.

Prosoma (Fig. 3). Dorsum brown with yellow median and lateral bands, marginal bands brownish grey, more or less broken into blotches. Margin black. Clypeus yellowish, with white pubescence. Chelicerae yellowish with greyish brown longitudinal stripes. Sternum yellowish, slightly greyish brown.

Eyes. Width of row I 42, row II 61, row III 77, row II-III 60. Diameter of AME 10, ALE 8, PME 24, PLE 18. Distance between AME 6, between AME and ALE 2.

Opisthosoma (Fig. 3). Dorsum with yellowish-grey lanceolate stripe bordered with few dark dots. Median light yellowish band behind lanceolate stripe with row of pairwise arranged dark dots, situated at each side of midline, continuing backwards. Dorsum on each side of light median band more or less speckled with black. Sides and venter yellowish, sides with few dark dots.

Legs (Table 1). Light brownish-yellow without annulation. TiI with two retrolateral spines.

Palp (Fig. 10, 44-49). Pt 0.50, Ti 0.50, Cy 1.15. Most of Fe, entire Ti and most of Cy greyish-brown. Apical part of Fe, most of Pt and Cy apically yellowish. All segments with dark hairs; yellowish parts of Fe and Pt in addition with whitish hairs (most of which missing in males examined; cf. Gravely's statement cited in Remarks below). Bulbus at level of embolus comparatively wide. Tegular apophysis with two acute projections in basal half (Figs 48, 49, cf. D. atropalpis : Fig. 13). Posterior subpaleal process large (Figs 45- 47), only anterior portion (portion above dotted line in Fig. 46) sclerotised, posterior portion membraneous, with pointed process partly surrounding paleal apophysis (Fig. 46). Embolus with laminar extension along inner side terminating in a process distally (Figs 44, 45).

Female (from Ootacamund, India). Total length 4.80, carapace 2.55 long, 1.90 wide.

Prosoma and opisthosoma (Fig. 4). Similar to male in colouration. Dorsal light yellowish median band on opisthosoma, with row of black dots in pairs posterior to lanceolate stripe, more distinct than in male because sides laterad to median band more dark.

Legs (Table 1). As in male but sometimes with very weak indistinct darker annulation.

Eyes. Width of row I 41, row II 60, row III 76, row II-III 60. Diameter of AME 10, ALE 8, PME 23, PLE 18. Distance between AME 5, between AME and ALE 2.

Epigyne (Figs. 21, 22, 25, cleared Fig. 31). Opening of median cavity narrow, most of it covered by a median septum. Each side with lateral elevation extending into a more or less triangle-shaped sclerite covering the receptacle. Bottom of median cavity corrugated, with characteristic shape in dorsal view (Fig. 22).

Size variation. Carapace length: males 2.20-2.55 (n=4), females 2.15-2.80 (n=15); tibia I vs. carapace length in Fig. 50.

Remarks. Gravely (1924) neither illustrated nor documented any particular details of the male apart from mentioning (p. 610) that "the male can be distinguished by the purer white pile on the upper surface of the femora of the palps, and by the continuance of this pile on to the inner side of the patella".

Tikander and Malhotra (1980) placed Pardosa lahorensis Dyal , 1935 (from Pakistan) as a synonym to P oakleyi but this was not considered in Mathew et al. (2009) or listed in Platnick (2010). I support this synonymy from studying material identified as Pardosa lahorensis collected in NW India (Ludhiana), only about 150 km from Lahore. Pardosa oakleyi has repeatedly (e. g., Mushtaq et al. 2003) been reported as a common species in fields of various crops in Pakistan.

Sadana (1971) illustrated an inflated bulbus, which he ascribed to P. lahorensis . Although this illustration is somewhat crude, it depicts a tegular apophysis (named 'conductor'), which is characteristic in shape as for P. oakleyi , as well as a subpaleal sclerite (named 'terminal apophysis').

Distribution. Pakistan, India, Bangladesh. A record from Vietnam (Pham Din et al. 2007) could not be verified as part of this study.