Orthomorpha glandulosa (Attems, 1937)

Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak, 2011, Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 131, pp. 1-161 : 60-62

Orthomorpha glandulosa (Attems, 1937) Figs 5152

Pratinus glandulosus Attems 1937: 119 (D).

Pratinus glandulosus - Attems 1938: 220 (D).

Orthomorpha glandulosa - Jeekel 1963: 265 (M); 1964: 361 (M, D); 1968: 56 (M); Hoffman 1977: 700 (M); Golovatch 1998: 42 (D, M); Enghoff et al. 2004: 38 (M, R).

Lectotype.

♂ (NHMW-3506), Vietnam, Hon Ba Island, Nhatrang, 06.1930, leg. C. Dawydoff.

Paralectotypes.

1 ♀ (NHMW-3506), same locality, together with lectotype. 1 ♀ (NHMW-3505), Vietnam, Darlac, frontier du Cambodge, 07.1930, leg. C. Dawydoff.

Lectotype designation proposed herewith is necessary to ensure the species is based on a complete male.

Redescription.

Length ca 38 mm (lectotype), 31-34 mm (♀), width of midbody pro- and metazona 3.4 and 5.0 mm (lectotype), 2.9-3.4 and 4.2-4.4 mm (♀), respectively (vs 3.0 and 5.0 in width, as given in the available descriptions ( Attems 1937, 1938)). Coloration of alcohol material upon long-term preservation dark grey-brown (Fig. 51) with contrasting pallid paraterga and epiproct, and light brown venter and legs (vs dark castaneous brown with paraterga and epiproct yellow, and venter and legs light red-brown, as given in the descriptions ( Attems 1937, 1938)).

Head usual, clypeolabral region densely setose, surface of vertex smooth, with a few setae flanking a distinct epicranial suture. Antennae long and slender (Fig. 51A, B & J), extending behind segment 4 (♂) or surpassing segment 3 (♀) dorsally. Head in width <collum <segment 2 <3 = 4 <5-16 (♂), or head <collum <segments 3 and 4 <2 <5-16 (♀), gently and gradually tapering thereafter. Collum smooth, with three transverse rows of setae traceable only as insertion points, 4+4 anterior, 2+2 intermediate, and 3+3 posterior setae; caudal corner of paraterga acutangular (ca 75°), nearly pointed (Fig. 51A, B & J). Tegument poorly shining; metaterga coriaceous, rugulose, each postcollum one with two rows of fully abraded setae borne on minute tubercles growing increasingly strongly reduced towards epiproct: 2+2 in front row and 3+3 in caudal one; prozona very finely shagreened, surface below paraterga finely microgranulate. Axial line rather evident, starting from collum. Paraterga very strongly developed (Fig. 51A-G & J-L), set high (at ca 1/4 metazonital height), in ♂ evidently upturned, lying above dorsum on postcollum segments, in ♀ mostly below dorsum, rather thin in lateral view, a little thicker on pore-bearing segments, on postcollum segments extending increasingly beyond rear tergal margin, better so in ♂, nearly pointed to pointed, caudal tip on paraterga 16-19 evidently curved mesad. Calluses on paraterga 2 delimited by a sulcus only dorsally, on following paraterga both dorsally and ventrally, rather broad. Paraterga 2 broad, anterior edge angulate, lateral edge with two minute incisions in anterior 1/3; posterior edge evidently concave (Fig. 51A, B & J). Paraterga 3 and 4 subequal, like subsequent paraterga, anterior edge broadly rounded, bordered and fused to callus, lateral edge with one minute incision in front 1/3. Ozopores evident, lateral, lying inside an ovoid groove, placed at about 1/3 metazonital length. Transverse sulcus complete on metaterga 4-18, incomplete on metaterga 2 and 3 (♂), or incomplete on metatergum 4 and complete on metaterga 5-18 (♀), shallow, not reaching bases of paraterga, ribbed at bottom, slightly sinuate anteromedially (Fig. 51A, C, F & J-L). Stricture between pro- and metazona narrow, shallow, beaded at bottom down to base of paraterga. Pleurosternal carinae complete crests only on segments 2-4 (♂, ♀) (Fig. 51B, D & E), each with an evident sharp denticle caudally, thereafter increasingly strongly reduced until segment 10 (♂, ♀). Epiproct (Fig. 51E-G & L) conical, flattened dorsoventrally, apical papillae small, dentiform, directed caudoventrally; tip subtruncate; pre-apical papillae small, lying close to tip. Hypoproct (Fig. 51G) roundly subtrapeziform, setiferous knobs at caudal margin small and well-separated.

Sterna sparsely setose, without modifications, but with a large, central, setose cone between ♂ coxae 4 (Fig. 51H & I). A paramedian pair of tubercles in front of gonopod aperture absent. Legs long and slender, midbody ones ca 1.3-1.4 (♂) or 1.2-1.3 (♀) as long as body height, prefemora without modifications, tarsal brushes present until ♂ legs 5.

Gonopods (Fig. 52) simple. Coxa long and slender, with several setae distodorsally. Prefemur rather large, densely setose, more than 2 times shorter than femorite + “postfemoral” part. Femorite very slender, evidently curved, not enlarged distad, “postfemoral” part demarcated by an oblique lateral sulcus; tip of solenophore small, trifid, with two subequal denticles (terminal and middle) and a larger subterminal lobule.