Calendula suffruticosa subsp. dersana A.C. Gonç. & P. Silveira, 2023
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publication ID |
https://doi.org/ 10.11646/phytotaxa.605.1.1 |
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persistent identifier |
https://treatment.plazi.org/id/E82AFC0A-1562-CA43-66ED-FF588554FA95 |
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treatment provided by |
Plazi |
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scientific name |
Calendula suffruticosa subsp. dersana A.C. Gonç. & P. Silveira |
| status |
subsp. nov. |
11b. Calendula suffruticosa subsp. dersana A.C. Gonç. & P. Silveira , subsp . nov. Type:— MOROCCO: Tetouan ,
Jbel Dersa, 468 m [35°35’26’’ N, 5°22’45’’ W], 7 June 2012, P GoogleMaps . Silveira & A.C.R.S. Gonçalves 3262 (holotype RAB!;
isotypes AVE!, MARK!).
Diagnosis:— Calendula suffruticosa subsp. dersana differs from C. suffruticosa subsp. suffruticosa by the predominantly white-arachnoid indumentum, instead of predominantly ± glabrous to glandulose, and cymbiform achenes length (5.5) 5.8–6.6 mm, instead of (8.8)
10.3–11.5 (14.5) mm. It differs from subsp. hosmarensis by having smaller rostrate achenes (8–10 mm long instead of 11–17 mm)
with dorsal spines, and its middle achenes being exclusively cymbiform (while subsp. hosmarensis has other types of achenes).
Description:––Perennial herbs. Stems (11) 17–32.5 (65) cm long, prostrate to decumbent, with white–arachnoid pubescence. Basal leaves (3.2) 3.5–7 (7.8) × (0.7) 1.1–2 (2.4) cm, (0.3) 0.4–0.4 (0.6) mm thick, oblanceolate to spatulate, apex obtuse, or sometimes ± acute, margins sub-entire to sinuate-dentate, with white–arachnoid pubescence. Capitula solitary, (1.8) 2.8–3.2 (3.6) cm diameter. Outer achenes rostrate (5) 6–10.2 (13.5) × (1.2) 1.3–1.5 (1.7) mm, straight or slightly curved, without dorsal spines, sometimes with 1–2 teeth, at the base or at the apex; middle achenes cymbiform (5.5) 5.8–6.6 × (3.6) 3.8–5.6 mm; inner achenes vermiculate-exalate (3) 3.3–3.5 (3.7) × (1.3) 1.5–1.7 mm, falcate to hemicyclic. Figure 17 View FIGURE 17 .
Habitat and distribution:—The species occurs on limestone rocks/cliffs, at an elevation of ± 500 m, and it is found in perhumid Mediterranean bio-climate. Endemic to Jbel Dersa in Tetouan ( Morocco).
Conservation status:—Critically Endangered (CR). This taxon is under numerous threats, especially due to agricultural intensification (grazing), climate change (extreme weather), and human intervention. The subspecies is rare and local with a narrowly restricted range; only one population is known close to abandoned pastures/fields. The number of mature individuals is estimated to be <50 based on field observations. The AOO is 8 km 2, which qualifies the taxon for CR. Therefore, we propose its conservation status as CR B2ab (ii, iii).
Chromosome number:—2 n = 32.
Genome size:—3.00 ± 0.07 pg.
Eponymy:—For Jbel Dersa in Morocco where it occurs.
Additional collections:— MOROCCO. Tanger-Tetouan-Al Hoceïma : Tetouan Jbel Dersa, [35°35’38” N, 5°23’06” W], 10 May 1921, C GoogleMaps . Pau s.n. ( MA, BC!); Tetouan Dersa, [35°35’38” N, 5°23’06” W], 13 April 1929, J GoogleMaps . Mas y Guindal s.n. ( MA!); Tetouan Jbel Dersa , [35°35’38” N, 5°23’06” W], 15 May 1930, P GoogleMaps . Font Quer ( BC!); Tetouan Jbel Dersa, 468 m [35°35’26” N, 5°22’45” W], 7 June 2012, P GoogleMaps . Silveira & A. C. R. S . Gonçalves 3262 ( AVE!); Tetouan , Jbel Dersa supra Semsa, [35°35’38” N, 5°23’06” W], 5 April 1911, C. J GoogleMaps . Pitard 1029 ( G!); Tetouan, Jbel Dersa, [35°35’38” N, 5°23’06” W], 30 March 1911, C. J GoogleMaps . Pitard 1027 ( G!); Tanger, Semsa, [35°35’37” N, 5°24’25” W], March 1930, J GoogleMaps . Mas y Guindal s.n. ( MA!); Tanger , Tetouan, Semsa, [35°35’37” N, 5°24’25” W], 5 April 1911, C. J GoogleMaps . Pitard 1028 ( G!) .
| P |
Museum National d' Histoire Naturelle, Paris (MNHN) - Vascular Plants |
| RAB |
Institut Scientifique |
| C |
University of Copenhagen |
| MA |
Real Jardín Botánico |
| BC |
Institut Botànic de Barcelona |
| J |
University of the Witwatersrand |
| A |
Harvard University - Arnold Arboretum |
| R |
Departamento de Geologia, Universidad de Chile |
| S |
Department of Botany, Swedish Museum of Natural History |
| AVE |
Universidade de Aveiro |
| G |
Conservatoire et Jardin botaniques de la Ville de Genève |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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