Tylopus bispinosus, Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010

Likhitrakarn, Natdanai, Golovatch, Sergei I., Prateepasen, Rujiporn & Panha, Somsak, 2010, Review of the genus Tylopus Jeekel, 1968, with descriptions of five new species from Thailand (Diplopoda, Polydesmida, Paradoxosomatidae), ZooKeys 72, pp. 23-68 : 26-27

Tylopus bispinosus sp. n. Figs 13

Holotype

♂ (CUMZ), Thailand, Tak Province, Umphang District, near Umphang City, ca 490 m, 16°2'20N, 98°52E, 6.07.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.

Paratypes:

1 ♂, 1 ♀, 2 juv. (CUMZ), same locality, together with holotype. 6 ♂, 4 ♀ (CUMZ), 3 ♂ (ZMUC), 3 ♂ (ZMUM), Tak Province, Umphang District, Doi Hua Mod, 900 m, 16°3'14N, 98°49'16E, 5.06.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn. 6 ♂, 1 ♀ (CUMZ), same Province, same District, Cave Ta Ko Bi, ca 530 m, 16°03'14N, 98°49'14E, 5.07.2009, leg. S. Panha, J. Sutcharit & N. Likhitrakarn.

Name:

To emphasize the spiniform processes h and z of the gonopod.

Diagnosis:

Differs from congeners in both processes h and z of the gonopod being spiniform.

Description:

Length 26 mm (holotype), 25-29 mm (♂), 33-38 mm (♀), width of midbody pro- and metazona 2.0 and 2.9 mm (holotype), 1.8-2.4 and 3.1-3.2 mm (♂), 2.4-2.7 and 3.3-3.8 mm (♀), respectively. Coloration of live animals black - brown (Fig. 1A): calluses of paraterga, venter and legs only slightly lighter, dark brown, but turning light brown in alcohol (Fig. 1 A–K).

Clypeolabral region of head very densely, vertigial region sparsely setose. Epicranial suture distinct. Antennae long and slender, reaching behind segment 4 (♂) or 3 (♀) dorsally. In width, head <collum <segments 3-4 <2 <5-16 (♂), or head = segment 3 <4 <collum <segments 5-16 (♀); thereafter body gradually and gently tapering towards telson (Fig. 1B).

Tegument generally rather smooth and shining, but prozona very finely rugulose, metaterga often rugose (Fig. 1 B–G); surface below paraterga finely microgranular (Fig. 1E, F). Collum with three transverse rows of setae: 5+5 in anterior, 2+2 in middle, and 4+4 in posterior row; paraterga evident, rounded, flap-shaped (Fig. 1B, C). Metaterga with two transverse rows of rather long setae: 2+2 in anterior and 2(3)+2(3) in posterior row, the latter often abraded, but then readily traceable as insertion points. Axial line at most barely visible only on metaterga. Paraterga strongly developed (Fig. 1 A–G), lying high (at 1/3-1/4 midbody height), only slightly inclined laterally, pointed caudally and acutangular already from segment 2, especially strongly so on caudal segments; calluses very thin on poreless segments, slightly thicker on pore-bearing ones; anterior 1/3 of poreless calluses with two evident (anterior larger, posterior smaller), lateral, setigerous incisions, but with only a single strong one (anterior) on pore-bearing calluses (Fig. 1 B–G); paraterga more strongly developed in ♂. Ozopores entirely lateral, lying in an ovoid groove about 1/3 in front of caudal corner, the latter always surpassing rear tergal contour (Fig. 1 B–H). Transverse sulcus evident on metaterga 4-18, reaching base of paraterga, evident and rather deep, faintly rugulose at bottom. Stricture between pro- and metazona very clearly ribbed (Fig. 1D). Epiproct tip faintly concave to subtruncate, pre-apical papillae evident (Fig. 1G). Hypoproct roundly subtrapeziform, caudal setae strongly separated (Fig. 1H). Pleurosternal carinae well-developed on segments 2-17 (♂) or 2-7 (♀), mostly as low bulges anteriorly and a distinct spine posteriorly (Fig. 1C, E, F).

Sterna moderately setose, without modifications; a deeply notched sternal lobe between ♂coxae 4 (Fig. 1I, J). Legs long and slender (Fig. 1B, C, H), especially so in ♂ compared to ♀ (1.3-1.5 versus 0.9-1.1 times as long as midbody height); prefemora distinctly bulged laterally (Fig. 1K), acropodites with particularly dense, nearly adpressed setae, including tarsal brushes.

Gonopods (Figs 2, 3) with lobe l well-demarcated; spine h very small; spine z considerably more prominent.