Scabiosa lucida barbata Nyar ., Enum. Pl. Vasc. Cheia Turzii: 280 (1939)

Novikov, Andriy, 2023, An annotated nomenclatural checklist of endemic vascular plants distributed in the Ukrainian Carpathians, Biodiversity Data Journal 11, pp. 103921-103921 : 103921

Scabiosa lucida barbata Nyar., Enum. Pl. Vasc. Cheia Turzii: 280 (1939)

Scabiosa barbata ≡ Scabiosa barbata Nyár. ex Chopyk & Fedoronchuk, Fl. Ukr. Carpath.: 436 (2015) [des. et nom. invalid.] *

Scabiosa pseudobanatica ≡ Scabiosa pseudobanatica subsp. barbata ( Nyár.) Chrtek, Preslia 57: 201 (1985); POWO: https://powo.science.kew.org/taxon/3007691-4

Asterocephalus lucidus = Asterocephalus lucidus [unranked] Asterocephalus lucidus a. alpicolus Schur, Enum. Pl. Transssilv.: 300 (1866); BHL: https://www.biodiversitylibrary.org/item/7364#page/320

Asterocephalus lucidus = Asterocephalus lucidus [unranked] b. subalpinus Schur, Enum. Pl. Transssilv.: 300 (1866); BHL: https://www.biodiversitylibrary.org/item/7364#page/320

Scabiosa columbaria = Scabiosa columbaria subsp. subalpina Brügger, Flora Curiensis. Naturgeschichtliche Beiträge zur Kenntniss der Umgebungen von Chur: 65 (1874) [nom. nudum]

Scabiosa columbaria = Scabiosa columbaria subsp. subalpina ( Brügger) Killias, Jahresber. Naturf. Ges. Graubündens 31: 82 (1887-1888); BHL: https://www.biodiversitylibrary.org/item/43011#page/278

Scabiosa columbaria = Scabiosa columbaria subsp. lucida var. subalpina ( Brügger) Braun-Blanq., Jahresber. Naturf. Ges. Graubünd. 58: 94 (1918); GBIF: https://www.gbif.org/species/12141628

Scabiosa lucida = Scabiosa lucida subsp. barbata f. alpicola (Schur) Prodan, Fl. Rep. Pop. Rom. 8: 685 (1961); POWO: https://powo.science.kew.org/taxon/3007684-4

Scabiosa lucida = Scabiosa lucida subsp. barbata f. hirticaulis ( Nyár.) Prodan, Fl. Rep. Pop. Rom. 8: 685 (1961)

Scabiosa lucida = Scabiosa lucida subsp. barbata f. perramosa ( Nyár.) Prodan, Fl. Rep. Pop. Rom. 8: 685 (1961)

Scabiosa lucida = Scabiosa lucida subsp. barbata f. subalpina (Schur) Prodan, Fl. Rep. Pop. Rom. 8: 685 (1961); POWO: https://powo.science.kew.org/taxon/3007687-4

Scabiosa lucida = Scabiosa lucida f. elata Nyár., Enum. Pl. Vasc. Cheia Turzii: 280 (1939); POWO: https://powo.science.kew.org/taxon/3007682-4

Scabiosa lucida = Scabiosa lucida f. hirticaulis Nyár., Enum. Pl. Vasc. Cheia Turzii: 280 (1939); POWO: https://powo.science.kew.org/taxon/3007681-4

Scabiosa lucida = Scabiosa lucida f. perramosa Nyár., Enum. Pl. Vasc. Cheia Turzii: 280 (1939); POWO: https://powo.science.kew.org/taxon/3007683-4

Scabiosa lucida = Scabiosa lucida f. scaposa Nyár., Enum. Pl. Vasc. Cheia Turzii: 280 (1939); POWO: https://powo.science.kew.org/taxon/3007680-4

Scabiosa lucida = Scabiosa lucida var. subalpina ( Brügger) Hayek & Hegi, Ill. Fl. Mitt.-Eur. 6(1): 308 (1908)

Scabiosa opaca = Scabiosa opaca Klokov, Novosti Sist. Vyssh. Nizsh. Rast.: 112 (1974) *; CoL: https://www.catalogueoflife.org/data/taxon/6Y55F; GBIF: https://www.gbif.org/species/4103554; IPNI: https://www.ipni.org/n/1004941-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0001046436; POWO: https://powo.science.kew.org/taxon/1004941-1

Scabiosa subalpina = Scabiosa subalpina Brügger, Jahresber. Nat. Gesell. Graubünd. 29: 137 (1886); GBIF: https://www.gbif.org/species/4104181; IPNI: https://www.ipni.org/n/320160-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000500648; POWO: https://powo.science.kew.org/taxon/320160-1; BHL: https://www.biodiversitylibrary.org/page/11661731#page/167

Scabiosa lucida - Scabiosa lucida Vill., Prosp. Hist. Pl. Dauphiné: 18 (1779) [p. p., tantum quod plantas ucrain. carpat.], non W.T.Aiton *; GBIF: https://www.gbif.org/species/7475424; IPNI: https://www.ipni.org/n/319978-1; WFO: http://www.worldfloraonline.org/taxon/wfo-0000500124; POWO: https://powo.science.kew.org/taxon/319978-1

Scabiosa lucida - Scabiosa lucida subsp. lucida Vill., Prosp. Hist. Pl. Dauphiné: 18 (1779) sensu Tasenkevych [non sensu orig.]

Conservation status

In Ukraine - LC ( Onyshchenko et al. 2022).

Distribution

SE Carpathian endemic.

Notes

There are four (https://powo.science.kew.org/taxon/319978-1, accessed on 05.06.2023) to five (https://europlusmed.org/cdm_dataportal/taxon/e6eec090-a11f-40b2-a8d1-a9f47863d58f, accessed on 05.06.2023) accepted subspecies of S. lucida Vill. - subsp. lucida (non-endemic taxon distributed in European mountains, starting from ca. 1000 m elevation - Štěpánek and Holub (1997)), subsp. Scabiosa lucida calcicola Błoński (endemic of W Carpathians - Štěpánek and Holub (1997), Danihelka et al. (2012)), subsp. Scabiosa lucida pseudobanatica (Schur) Holub (problematic taxon with Carpathian-Pannonian distribution range that prefers lower elevations and occurs in Ukraine, Slovakia and Romania, but often overlooked or confused - Chrtek (1985), Chrtek and Goliašová (1985), Tasenkevich (2006)), subsp. Scabiosa lucida stricta (Waldst. & Kit.) Jasiewicz (distributed in southern Europe) and subsp. Scabiosa lucida barbata Nyár. (south-eastern Carpathian endemic - Kliment et al. (2016)).

In the Flora of UkrSSR ( Kotov 1961: p. 378), S. lucida is indicated as having var. subalpina ( Brügger) Hegi with S. subalpina Brügger, Zur. Fl. Tirol (1860) amongst its synonyms. However, POWO (https://powo.science.kew.org/taxon/77227194-1, accessed on 05.06.2023), WFO (https://list.worldfloraonline.org/wfo-0000500277, accessed on 05.06.2023) and Worldplants (https://www.worldplants.de/world-plants-complete-list/complete-plant-list?name=Scabiosa-columbaria, accessed on 07.06.2023) indicate S. subalpina to be a synonym for S. columbaria L., a quite distinct species. After checking, it was found that Brügger did not describe S. subalpina in his mentioned work "Zur Flora Tirols" ( Brügger 1860). IPNI (https://www.ipni.org/n/320160-1, accessed on 05.06.2023) says that Brügger described this species much later, in 1887-1888 (i.e. S. subalpina Brügger, Jahresber. Naturf. Ges. Graubündens 31, Beil. 82 (1887-88). However, this is also wrong because IPNI references the work of Killias (1887), who only repeated an epithet Scabiosa subalpina for S. columbaria subsp. subalpina (the rank of subspecies is indicated by the author in the taxon’s description) following Brügger (1874), who, in turn, published this subspecies without the protologue. Considering the mistake in IPNI, it is worth noting that complete and correct nomenclatural citations for this subspecies should be S. columbaria subsp. subalpina Brügger, Fl. Cur.: 65 (1874) [nom. nudum] and S. columbaria subsp. subalpina ( Brügger) Killias, Jahresber. Naturf. Ges. Graubündens 31: 82 (1887-1888). Nevertheless, this does not answer the question about where the initial Brügger’s species name has been applied. In fact, Brügger first applied the epithet Scabiosa subalpina in 1874 (without any protologue provided) and later, in 1886, with a subsequent self-reprint of the last paper in the same 1886 that, however, has changed pagination ( Brügger 1874, Brügger 1886a, Brügger 1886b). Therefore the correct nomenclatural citation for this species should be S. subalpina Brügger, Fl. Cur.: 65 (1874) [nom. nudum] et Jahresber. Nat. Gesell. Graubünd.: 137 (1886). Brügger (1886a) and Brügger (1886b) described S. subalpina as an intermediate species between S. columbaria and S. lucida. It was Hayek and Hegi (1908), who, on page 308, determined S. subalpina belonging to S. lucida and applied the combination S. lucida var. subalpina ( Brügger) Hayek & Hegi, Ill. Fl. Mitt.-Eur. 6(1): 308 (1908). Schur (1866) also believed that Scabiosa subalpina plants belong to the Scabiosa lucida group and applied the new name Asterocephalus lucidus [unranked] b. subalpinus Schur, Enum. Pl. Transssilv.: 300 (1866). Later, Pawłowski (Planta Poloniae Exsiccata deposited at KW) synonymised S. lucida var. subalpina with S. lucida var. lucida Considering this, I believe that the mentioned databases mistakenly synonymise S. subalpina (and derivatives) with S. columbaria . Instead, S. subalpina should be considered as a synonym of S. lucida. Moreover, after herbarium inspection, I found that at least part of the specimens identified as S. lucida var. subalpina belong to S. lucida subsp. barbata .

It is important to note that Brügger mentioned many taxa (e.g. Hepatica rhaetica, Malus hortensis, Batrachium micranthum, Nasturtium Scabiosa montanum etc.) for the first in Flora Сuriensis ( Brügger 1874); however, this book is entirely ignored by IPNI and, consequently, many other databases, for some reason.