Ptomaphaminus fagei Perreau, 2009

Schilthuizen, Menno, Perreau, Michel & Njunjic, Iva, 2018, A review of the Cholevinae from the island of Borneo (Coleoptera, Leiodidae), ZooKeys 777, pp. 57-108 : 85

Ptomaphaminus fagei Perreau, 2009 Figures 12 q–r, 13 k–l

Ptomaphaminus fagei Perreau, 2009. Perreau (2009: 2, fig. 4); type from Gunung Marang, Kalimantan, Borneo (in CMPR).

Material.

Kalimantan Timur. Kebupaten Kutai Timur, karst of Mangkalihat, Mt Marang, Gua Sedepan, 8.vi.2002 (leg. M. Perreau, Expédition du Kalimanthrope, TXEX), 1 male, 1 female paratypes.

Description.

Length 1.50-1.90 mm. Colour light brown. Winged. Eyes with 15 ommatidia. Body very elongate, parallel-sided. Pronotum approximately 1.6 times wider than long. Elytra approximately 1.4 times longer than wide. Female sutural angle of elytra expanded backwards in a sharp tooth. Male protarsi 0.75 times as wide as the apex of protibia. Spiculum gastrale of the male genital segment expanded, with a triangular apical part (Figure 13k). Aedeagus 4.8 times as long as the body length, parallel-sided at the base, triangularly narrowed in the last third of the length in dorsal view (Figure 12q), thick at the base, abruptly thinned in the two apical third of its length and ending with a short ventrally deflexed hook in lateral view (Figure 12r). Five lateroventral lateral external setae and three more ventral central setae on each side. Internal stylus of the endophallus straight. Parameres with five apical external and three apical internal setae. Female genital segment with reduced gonocoxites, spermatheca bilobate, with helicoidal spermiduct, and without apical sclerotised plate (Figure 13l).

Differential diagnosis.

Among the species with similarly reduced eyes and/or spiniform female elytral apices, P. fagei is unique in having an aedeagus that shows an abrupt narrowing (in lateral view) in the apical third.

Habitat and distribution.

Known from two caves of Gunung Marang, Kalimantan, Indonesia: Gua Sedepan and Gua Gala.