Pyropteron leucomelaena leucomelaena ( Zeller, 1847 )

Bartsch, Daniel, Pühringer, Franz, Milla, Liz, Lingenhöle, Arthur & Kallies, Axel, 2021, A molecular phylogeny and revision of the genus Pyropteron Newman, 1832 (Lepidoptera, Sesiidae) reveals unexpected diversity and frequent hostplant switch as a driver of speciation, Zootaxa 4972 (1), pp. 1-75 : 39-40

Pyropteron leucomelaena leucomelaena ( Zeller, 1847)

( Figs 120–122 , 221, 222 , 229, 230 )

Sesia leucomelaena Zeller, 1847 : Isis, 40 (1): 12. Lectotype ♂: Macri [ Turkey, Fethiye] (NHMUK).

Sesia therevaeformis Lederer, 1853 : Verhandlungen des zoologisch-botanischen Vereins in Wien, Abhandlungen 2 [“1852”], 2: 82. Syntypes: 2♂ (?), 4♀, Spalato [ Croatia, Split] and Brussa [ Turkey, Bursa] (lost? Not in MFNB).

Sesia cretica Rebel, 1916 : Annalen des k.k. Naturhistorischen Hofmuseums, 30: 143. Syntypes: 2♀, Greece, Crete, Kristallenia and Mallaes (NHMW).

Chamaesphecia leucomelaena f. cyrnea Le Cerf, 1920 : Études de Lépidoptérologie Comparée , 17: 506. Syntypes: 15♂, 5♀, France, Corsica (MNHN).

Distribution. The typical subspecies is known from southern France in the west across southern Europe to Turkey in the east. It also occurs on the Mediterranean islands of Corsica, Sardinia, Sicily, Crete and Cyprus ( Laštůvka & Laštůvka 1995, 2001, Špatenka et al. 1999). Based on Gorbunov (pers. communication), this species is absent from Crimea. Specimens from central and northern Spain are here provisionally assigned to the typical subspecies; however, more detailed analyses are needed to map the precise distribution of the two subspecies of P. leucomelaena in the Iberian Peninsula.

Specimens examined. 17♂, Spain, Huesca, Graus, 30. VI .–10. VII .2009, leg. Riefenstahl & Wagenblass ( Bartsch gen. prep. 2020-04) ( Fig. 222 ) ; 2♂, Spain, Teruel, Frias / Albarracin , 1400m, 1.–10. VII .1999, leg. Riefenstahl & Wagenblass (SMNS); 1♂, Spain, Castillia-La-Mancha, Umgeb. Los Alares, 26. VI . 2012, 580m, leg. AL; 1♀, same locality, 19. VI . 2012, 580m, leg. AL; 1♂, Spain, Castilla-La-Mancha, Horcaja de los Montes, 435m, 18. VI .2012, leg. AL; 1♀, same locality, 19. VI .2012, leg. AL (CAL); 6♂, Spain, Teruel, Portillo de Guadalaviar (El Portillo), 1760–1770 m, 9. VII .2018, leg. FP (CFP); 1♂, Spain, Teruel, Albarracin , 1200m, 30. VI .1993, leg. DB; 1♂, Spain, Huesca, Graus , 500m, 30. VI .2009, leg. AL ( Bartsch gen. prep. 2019-52) ( CDB) ; 1♂, Spain, Teruel (Aragon), Albarracin, 25. VI .1928, leg. Predota; 1♂, Spain, Catalonia, Vidreras nr. Gerona, 12. VI .1993, leg. A. Pürstinger; 1♂, same data, 15. VI .1993 ( CFP) . 1♂, France, Ardeche , Saint-Maurice-d’Ibie, 350 m, 6. VII .1998, leg. B. Vanholder (CCDB-02110 B07, gen. prep. FP 98/64); 1♂, France, Corsica, Castirla NW Corte, 950 m, 10. VI .1999, leg. H. Fischer (BOX-2219 H11, gen. prep. FP 99/11) (CFP); 1♀, France, Alpes de Haut Provence , Luberon, Manosque, 14. VI .2006, leg. E. Blum ( Bartsch gen. prep. 2019-53) ( Fig. 230 ) ( CDB) . 5♂, Provence, Montbrun les Bains , 6., 7. and 8. VII .1991, leg. DB; 1♀, France, Drome , Séderon, Col de la Pigiere, 25. VII .1992, leg. DB; 1♂, France, Provence , Esterel, Agay, 25. VI .1991, leg. DB (CDB). 3♂, Italy, Sicily, PA, N Castelbuono, 120 m, 10. VI .2001, leg. FP (DNAtax 02-18, gen. prep. FP 02/08); 12♂, Italy, Sicily, SR, Pedagaggi NW Sortino, 700 m, 15. VI .2001, leg. FP (DNAtax 02-19, gen. prep. FP 02/06); 2♂, Italy, Sicily, SR, W Sortino, 530 m, 13. VI .2001, leg. FP; 2♂, ibid., 600 m, 15. VI .2001, leg. FP (BOX-2219 H12, gen. prep. FP 02/09) (CFP); 1♂, Italy, Sicily, W of Sortino , 600m, 15. VI .2001, leg. FP; 1♂, Italy, Abruzzi, Tuffo , 800m, 22. VII .1991, leg. HR (CRS); 1♂, Albania, Lezha, W Reps, 260 m, 8. VII .2016, leg. FP; 1♂, Bulgaria, Burgas, Chernomorets , 30 m, 9. VII .2014, leg. FP; 5♂, Bulgaria, Slivno, N Sliven (Slivno), 390 m, 3. VII .2014, leg. FP; 2♂, ibid., 400 m, 10. VII .2014, leg. FP; 1♂, ibid., 25. VI .2016, leg. FP; 1♂, Greece, Crete, Chania, Marathi (Halbinsel Akrotiri), 10 m, 6. VI .2000, leg. FP; 1♂, ibid., 50 m, 7. VI .2000, leg. FP (CCDB-02113 F04); 2♂, Greece, Crete, Rethymnon, Skaleta E Rethymnon, 2. VI .1996, leg. FP (photo 24/1/40-42, 2/35-41); 1♂, ibid., 4. VI .1996, leg. FP; 6♂, ibid., 6. VI .2000, leg. FP (CCDB-02110 B10, photo 41/6/1- 4); 2♂, ibid., 7. VI .1996, leg. FP; 1♂, Greece, Fthiotis , 5 km N Bralos, 450 m, 30. VI .2002, leg. FP (DNAtax 01-16); 3♂, ibid., 3. VII .2003, leg. FP (BOX-2221 F11); 1♂, Greece, Fthiotis, Iti (Oiti), 670 m, 22. VI .2002, leg. FP (photo 51/5/33-41); 1♂, Greece, Fthiotis, Pavliani , 1000 m, 22. VI .2002, leg. FP; 5♂, Greece, Ioannina, Kipi (= Kipoi , Timfi Mts ), 780 m, 28. VI .2002, leg. FP (DNAtax 02502, photo 52/5/11-13); 3♂, Greece, Ioannina, Koukouli , 930 m, 30. VI .2003, leg. FP (CCDB-02110 B08); 1♂, Greece, Komotini , N Drama, 350 m, 6. VII .2003, leg. FP; 1♂, Greece, Larissa, Ossa Mts , SW Stomio (n), 1000–1090 m, 20. VI .2002, leg. FP; 5♂, ibid., 2. VII .2003, leg. FP (BOX- 2220 A05) (CFP); 1♂, Greece, Peloponnese, inner Mani , Kendro, 1200–1800m, 7. VII .2005, leg. DB; 1♀ ( Fig. 122 ), Greece, Peloponnese, outer Mani, Kariovouni , 600m, e.l. ex. Sanguisorba minor , larva 19.IX.2000 adult 30. V .2001, leg. DB (CDB); 1♂, Bulgaria, Vlas, 27. VI .–3. VII .2005, leg. Riefenstahl; 1♂, Turkey, Antalya, Saklikent , 640m, 28. VI .2002, leg. Riefenstahl; 2♂, ibid., 1380m, 28. VI .2002, leg. Riefenstahl; 9♂, ibid., 900m, 28.–29. VI .2002, leg. Riefenstahl; 2♂, ibid., 1610m, 29.–30. VI .2002, leg. Riefenstahl; 10♂, Turkey, Anatolia, Feke , 1000m, 5. VII .1992, leg. Riefenstahl (SMNS); 7♂, Turkey, Toros Daglari, road Antalya-Saklikent , 1000m, 29. and 30. VI .1998, leg. DB; 7♂ ( Figs 120–121 ), ibid., 600m, 16. VI . and 3. VII .2002, leg. DB; 1♂, Turkey, Mersin, Pusatli Dagi , 10 km S Kirobasi, 1260m, 29. VI .2002, leg. AL ( Bartsch gen. prep. 2019-50) ( Fig. 221 ) ; 9♂, ibid., 1280m, 29. VI .2002, leg. DB; 1♀, Turkey, Konya, Beysehir , 1170m, 27. VI .2002, leg. AL ( Bartsch gen. prep. 2019-51) ( Fig. 230 ) ; 2♂, Konya, Taskent , 1570m, 1. VII .2002, leg. DB; 2♂, Turkey, Konya, 45km S Beysehir, Bakaran , 1420m, 18. VI . and 2. VII .2002, leg. DB; 1♂, 1♀, Turkey, Adiyaman, 25km W Gölbasi , 24. VI . 1993, leg. G. & M. Novak; 2♂, Cyprus, Limassol, Souni , 400m, 19. V .2001, leg. G. & M. Novak; 1♂, Cyprus, Paphos, Akamas, Ag. Minas, 23. V .2001, leg. G. & M. Novak; 1♂, 2♀, Cyprus, Troodos , 2km NW Fioni, 1200m, larvae 25. III .2000, e.l. ex. P. minor , leg. DB (CDB).

Figures

FIGURES 120–134. Pyropteron species. 120–122. Pyropteron leucomelaena leucomelaena, (120–121) ♂, Turkey, Antalya, (122) ♀, Greece, Peloponnese. 123–128. P. leucomelaena blaesii ssp. nov., (123–124) ♂ Spain, Sierra Nevada, (125) ♀, (126– 127) ♂ holotype, Morocco, High Atlas, (128) ♀, Morocco, Middle Atlas. 129–131. Pyropteron aistleitneri, (129–130) ♂, (131) ♀, Spain, Granada. 132–134. Pyropteron kautzi, (132–133) ♂, Spain, Granada, (134) ♀ lectotype, Spain, Granada (image Michael Falkenberg). Scale 10mm.

FIGURES 221–225. Pyropteron species, male genitalia. 221–222. Pyropteron leucomelaena leucomelaena, ♂, (221) Turkey, Mersin, genitalia slide DB 2019-50, (222) Spain, Huesca, genitalia slide DB 2020-04. 223. P. leucomelaena blaesii ssp. nov., ♂, paratype, Morocco, Middle Atlas, genitalia slide DB 2020-11. 224. Pyropteron hellenicum sp. nov., ♂, paratype, Greece, genitalia slide DB 2019-40. 225. Pyropteron borreyi, ♂, Morocco, High Atlas, genitalia slide DB 2007-50. Scale 1mm.

FIGURES 226–232. Pyropteron species, female genitalia. 226. Pyropteron jordanicum sp. nov., ♀, Israel, slide DB 2020-06. 227–228. Pyropteron triannuliformis, ♀, (227) Jugoslawia [Croatia], Porec, slide DB 2019-55; (228) Greece, Pindos, slide DB 2019-42. 229–230. Pyropteron leucomelaena leucomelaena, ♀, (229) Turkey, Konya, slide DB 2019-51, (230) France, Luberon, slide DB 2019-53, 231. P. leucomelaena blaesii ssp. nov., ♀, Morocco, Middle Atlas, slide DB 2020-12. 232. Pyropteron borreyi, ♀, Morocco, High Atlas, slide DB 2007-51. Scale 1mm.