Promanodes, Kolibáč, Jiří, Schmied, Heiko, Wappler, Torsten & Kubisz, Daniel, 2010

Kolibáč, Jiří, Schmied, Heiko, Wappler, Torsten & Kubisz, Daniel, 2010, A description of Promanodes serafini gen. et sp. nov. from Baltic amber, with a review of related New Zealand Promanus Sharp, 1877 (Coleoptera: Trogossitidae), Zootaxa 2620, pp. 29-44 : 31-32

Promanodes gen. nov.

Type species. Promanodes serafini sp. nov.

Diagnosis. The new genus is very similar to the recent Promanus , the both genera share especially deep and incurvate frontoclypeal suture ( Figs 1 , 30–32 ), distinctly elevated eyes ( Figs 3, 5, 6 , 23–25 ), weakly or no way projecting anterior pronotal corners ( Figs 1, 2 , 23–25 ), elytra with weak or inconspicuous carinae widest at about 2/3 of length ( Figs 1 , 26–28 ), radial cell obliquely situated ( Figs 4 , 8 , 17 ), dorsal body surface sparsely pubescent or nearly bare ( Figs 1 , 23–25 ), femora conspicuously clavate ( Figs 2 , 22 ), very small trochanters ( Figs 2 , 22 , 35, 36 ), and especially abdomen with 6 visible sternites ( Figs 2 , 7 ; Kolibáč 2005).

The new genus is based on an apomorphic state of the prothorax (coxal cavities nearly closed; Fig. 7 ) while the securiform terminal joint of the maxillary palps ( Fig. 21 ; Kolibáč 2005) is an apomorphy of Promanus . The main differences are listed in the Table 1.

Remark. The extant Holarctic lophocaterine genus Grynocharis Thomson, 1859 is recently sympatric with Promanodes gen. nov. and was referred from Baltic amber as well as another lophocaterine genus Lophocateres Olliff, 1883 , with originally southeastern Asian distribution (one pest is a cosmopolitan). That is why, morphological differences of Grynocharis and Lophocateres are also added in Table 1 and in a character analysis ( Table 3, Fig. 37).

Etymology. The new genus-group name is a combination of Promanus and an adjectival derivative of eidos (Greek, meaning, “kind” or “having the form of”). The name is masculine (while generic words ending in the noun eidos are neuter, those ending in the adjectival derivatives, such as – odes or – oides, may be in practice any gender [masculine, feminine, or neuter]: Brown 1954).

in parentheses denote characters for Table 3.

Character Promanodes gen. nov. Promanus Grynocharis Lophocateres

Maxillary palpi (0) last joint coniform last joint securiform last joint coniform, last joint coniform truncate at apex

Labial palpi relatively long; last joint short; last joint nearly short; conical short; conical narrowly securiform cylindrical, weakly

dilated towards apex

Head dorsally bare with decumbent with fine decumbent with fine semi-erect pubescence pubescence pubescence

Figures

FIGURES 1 – 6. Promanodes serafini gen. et sp. nov.: (1) body dorsally; (2) body ventrally; (3) Pro- and mesothorax ventrally; (4) wing; (5) Head ventrally; µCT scans: (6 a) body ventrally; (6 b) body laterally. Scale bar 1 mm. (Photo T. Wappler / A. Bergmann)

FIGURES 23 – 32. Habitus photo: (23) Promanus auripilis Sharp; (24) P. subcostatus Broun; (25) P. depressus Sharp. Apex of elytra: (26) P. auripilis Sharp; (27) P. subcostatus Broun; (28) P. depressus Sharp. Antenna: (29) P. auripilis Sharp. Head dorsally: (30) P. auripilis Sharp; (31) P. subcostatus Broun; (32) P. depressus Sharp. (Photo J. Kolibáċ).

FIGURES 7 – 11. 7 – 9 Promanodes serafini gen. et sp. nov.: (7) body ventrally, partial reconstruction; (8) radial cell area of wing; (9) mouth parts ventrally. 10 – 11 Promanus subcostatus Broun: (10) female tergite VIII; (11) female sternite VIII. (J. Kolibáċ del.).

FIGURES 17 – 22. Promanus depressus Sharp: (17) wing; (18) female sternite VIII; (19) female tergite VIII; (20) maxillary palp; (21) labial palp; (22) hind leg. (J. Kolibáċ del.).

FIGURES 33 – 36. Head ventrally: (33) P. auripilis Sharp; (34) P. subcostatus Broun. Pro- and mesothorax ventrally: (35) P. auripilis Sharp; (36) P. subcostatus Broun. (Photo J. Kolibáċ).