Asterocheridae Giesbrecht, 1899
|
publication ID |
https://doi.org/ 10.12651/JSR.2024.13.2.185 |
|
persistent identifier |
https://treatment.plazi.org/id/ED1887CB-FFFF-FFEE-FCEE-B35FB4996348 |
|
treatment provided by |
Felipe |
|
scientific name |
Asterocheridae Giesbrecht, 1899 |
| status |
|
Family Asterocheridae Giesbrecht, 1899 View in CoL Genus Asterocheres Boeck, 1859
Remarks on the valid species. WoRMS ( WoRMS Editorial Board, 2023) has recognized 91 species as valid in the genus Asterocheres . We accept 88 of those 91 species as valid, except the following three species: A. berniciensis (Brady, 1899) ; A. carausi Marcus & Por, 1960 ; and A. ornatus ( Brady & Robertson, 1876) . Asterocheres berniciensis was described under the name of Cyclopicera berniciensis based on the male. The description and illustration of A. berniciensis exhibit no diagnostic feature of Asterocheres and we recognize it here as a species inquirenda. We follow Stock (1969) who treated A. carausi to be a synonym of A. uncinatus (Krichagin, 1873) . Asterocheres ornatus (originally as Ascomyzon ornatum ) is currently recognized as a species of the genus Acontiophorus Brady, 1880 ( Brady, 1880; Giesbrecht, 1899). WoRMS has recognized A. siphonatus Giesbrecht, 1897 as a synonym of Dermatomyzon nigripes ( Brady & Robertson, 1876) , but we treat it a distinct species of Asterocheres , as redescribed by Conradi & Bandera (2011) and in the present paper. Therefore, a total of 89 species in
Characters(1-17) Species Data source 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17
A
A
A
A
A
Bendera
.
.
.
.
. alter bacescui aesthetes antillensis astroidicola
Eiselt
& Lopez-Gonzalex
Marcus
Ho
Varela
, 1965
Conradi
, 1984
,
,
1965
2010
,
, 2006
3
1
1
3
4
1
3
3
1
- -
-
3
2
2
21
21
21
19
-
-
-
-
-
-
4
4
2
4
2
3
1
2
3
3
2
5
1
2
1
2
2
2
2
2
3
4
3
1
+
+ +
+ X X X
X
X X + + +
-
- 1
2
2 1 3 3 3
3
- 2
X
X 1 -, 3 17
18
18 18 - X X +
-
- Eiselt Varela Present
Marcus
Conradi ((1965 2010
(
study
1965
et al) b.
)
) (2006)
A
A
.
.
banderaae bahamensis
Kim
Kim
&
, 2010
Lee, 2023
1
1 1
1 3
3
19
20 -
- 3
3
1
1
2
2 1
2
2
4
X + + X X X + 1 2 3 3 1, 3 3 17 17 X X Kim Kim (& 2010 Lee) (2023)
A
Johnsson
. bimbarrensis
& Neves
Bispo
, 2006
, 3 2 3 20
-
- 3 3 3 2 -
+ X + 2 3 X - 18 - X - Bispo et al. (2006)
A
A
A.
.
. boecki bulbosus brevisurculus
(Brady
Malt
,
Kim
,
1880
1991
, 2005
) 1
3
1
1
2
3
3
3
2 20
21
19 -
- 3
2
4
1
3
2 1
1
4
1
2
1
2
2
4 + + - X
X
X X + +
123 1 2 123 3 3 X 1 17 18 X X Kim Malt Bandera ((2005 1991 &)) Conradi; Bandera (2016 & Mercedes) (2009)
A
A
A
A
A
A
.
.
.
.
.
. canui cubensis corneliae crenulatus complexus crinoidicola
Giesbrecht
Varela
Schirl
Johnsson
Stock
Humes
,
,
2010
,
,
1973
1960
1897
,
,
2000
1999
-
2
2
1
3
3 1
1
1
3
1
- -
2
2
2
2
2
20
21
19
21
19
-
-
-
-
-
-
-
3
2
4
3
4
3
3
1
1
2
-
3
2
3
2
2
2
2
2
1
2
2
2
2
4
4
3
-
-
+
+
+
+
+
+ X
X X
X
X + +
+
+
+
+
+ 1= 1 1
2 1
2 2 3 3
3
3
3
3
X
2
-
- - - 18 18 17
18
-
-
X
X
X X
-
-
Kim
Canu Stock Schirl
Varela
Johnsson
(2010
(((1892
(
1960 1973
2010
(1999
)
,))
a as;;
)
Bandera Bandera) A. lilljeborgi & & Conradi Conradi) ((2014 2016)); Schirl(1973)
A
A
A.
.
.
Kim
Giesbrecht
(
,
Norman
2016
,
, 1897
1868)
3
1
4 1
1
1
X
2
3 20
21
20
-
-
-
3
4 2
1
-
2
1
-
1
2
-
3
-
- +
+
+ X X - X + - 2 1 3 3 3 1 2 1 -, 3 18 17 - X X - Kim Bocquet Bandera Giesbrecht (2016 et &) (al Conradi 1899.(1963) (, 2009 as A. b violaceus );)
A
A
Lalana
.
.
ellisi espinosai
, 2007
Hamond
Varela
, 1968
, Ortiz &
3
2 1
2
2
3 21
21
-
-
-
4
3
1
3 2
5 2
2 3
2
- + +
X X + + 2 1 3 3 X - 18 - X - Varela Hamond, Ortiz (1968 &); Lalana Bandera (2007 & Mercedes) (2009)
A
A
Baviera
.
& Zaccone
Bandera
Crescenti
, 2010
&
,
Conradi, 2014
4
4 1
2
3
3 21
21
- -
4
4
1
3
4
3 1
2
3
X
+
X
X
+ + 1 1 3 3 X 1 - 17 18 - X X - Bandera Crescenti & et Conradi al. (2010 (2014))
A A
A
.
.
.
flustrae galeatus fernandezmilerai
Ivanenko
Kim, 2010 Varela
& Smurov
, 2010
, 1997 3
3 1 1
1
1 2 3
3
19 20
21
-
-
3
4
3 3 3
3 2
4 3
2
2 2 3 3
3
+
+ X X X X + + 3 2 1 3 2 1 X X 18 18 X + Kim Varela Ivanenko (2010 (2010 &) Smurov a) (1997)
A
. Lalana garridoi
X + 1 3 X 17 X Varela et al. (2007)
, 2007
Varela, Ortiz & 2 1 3 20 - 2 3 2 1
Continued.
Characters(1-17) Species Data source 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17
A
A
A
Lopez-Gonzalez
.
.
.
hoi hirsutus halichondriae
Bandera
Bandera
&
, Stock 2005
Conradi
Conradi
, 1966
, 2013
& 5
2
2 2
1
2
3
2
4
21
20
21 -
-
-
4
4
3 3
3
3
2
4
5
2
2
2
3
-
-
+ + X X X + + 1 1 1 3 3 3 X 2 - 17 18 - X X - Stock Bandera Bandera (1966 et & al Conradi);. Bandera (2005 () 2013 & Conradi) (2013)
A
A
A
A.
.
.
. humesi indicus horridus hongkongensis
Varela Sewell
Kim,
,,
2016
2012
Malt
1949
, 1991
1
3
3
4
2
2 1
2
- 2
2
3
19
21
19
20
-
- -
-
3 3
4
2
2
3
1
2
1
2
4
-
2
1 2
1
3
2 1
-
+
+
+
+
+ X
X X
X
+ +
+
-
?
2
2 2
1
3
3 3
3 2
X
-
-
, 3
17
18 -
- + -
-
-
Kim Malt
Varela Sewell
((2016 1991
((2012 1949
));
)
Bandera
); Bandera
&
&
Mercedes
Mercedes
(2009
(2009
)
)
A
A.. indivisus intermedius Kim (Hansen, 2010, 1923) 4 1 3 1 - 2 21 20 - - 4 3 1 - 2 5 1 - 3 - + - X -
+ - - 2 3 - 1, - 2 18 - X - Kim Hansen (2010 (1923)); Bandera & Conradi(2009a)
A
A
A
A
A
.
.
.
.
.
kimi latus lalanai lilljeborgi jeanyeatmanae
Varela
(Brady
Verela
Boeck
,
,
2012
1872
, 2013
Yeatman
, 1859
)
, 1970
2
5
2
4
5
1
3
1
3
1
-
4
4
2
2
20
20
21
21
19
-
-
-
-
-
2
4
3
3
4
1
1
3
3
3
2
3
2
3
3
2
2
2
1
1
4
3
3
2
-
+ +
+
+ + X
X X X
X
X + + + + 1 1 1 1 2 3 3 3 3 3 X X - - - 17 18 - - - X X - - - Kim Varela Varela Bandera Yeatman (2016 ((2012 2012 & (1970) Conradi), as) A. stocki (2009) b)
A. longisetosus Nair & Pillai, 1984 1 1 2 20 - 2 1 - 1 1 +
A
Conradi
& Lopez-Gonzalez
Bandera,
, 2007
2 1
-
3 21 -
-
4 3 2
-
1 3
+
-
X X
-
+ + 2 2 3 3 X
-
- 18 - X
-
- Nair Bandera & Pillai et al (. 1984 (2007))
A. major Thompson & A. Scott, 1903 2 3 20 4 3 2 4 X 3 1 17 Thompson & Scott(1903)
A
A
.
.
Scott, 1903
Thompson & 1 1 - 20 -
-
4 3 - 2 3
+ X - 1 3 -
- -
- -
-
Thompson & Scott(1903)
A
A.. micheli maxillatus (Gurney Stock,, 1927 1987) 1 3 1 3 - 1 20 18 - 3 3 - 2 4 2 2 3 - + - X - + - 2= - 3 1 - - - - Stock Gurney (1987 (1927))
A. minor Thompson & A. Scott, 1903 3 1 - 20 - 4 - - 2 - - - - 3 1 - 17 - Thompson & Scott(1903)
A. minutus (Claus, 1889) 4 1 1 21 - 4 1 2 1 3 - X - 3 1 1 17 X Bocquet et al. (1963);
Conradi & Bandera(2011)
A
Rocha
. neptunei
& Neves
Johnsson
, 2002
, In: Johnsson, 3 1 3 20 -
-
3 3 4 2 1
+ X + 1 3 - 17 X Johnsson et al. (2002) A
Suárez-Morales
. nidorellae Reyes-González
, 2021
& 1 1 3 21 4 2 2 2 3
+
X
+
1
=
2 3 3 18 X Reyes-González &
Suárez-Morales(2021)
A
A
A
.
.. oricurvus orientalis nudicoxus Kim
Sewell
Kim,, 2010 2010
, 1949
1 3
2
3 3
1
2
2
3 21 20
20 -
- - 3
4
4 1 1
-
2
3
5 2
2
2 3 1
-
+ + + X X X X X + 2 1 2 3 3 3 1, 1 - 2 18 18 - X X - Kim Kim Sewell ((2010 2010 (1949))
A A.. ovalis parvus Sewell Giesbrecht, 1949 , 1897 - 4 1 - - 3 - - - - 2 1 - 1 - 2 - 4 - + - X - + - 1 2 3 3 X - 14 - X - Sewell Giesbrecht (1949 (1899)); Bandera) & Mercedes (2009)
Continued.
Characters(1-17) Species Data source 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17
A
A
A
A
A
.
.
.
.
.
planus pilosus plumosus peniculatus proboscideus
Kim
Kim
Kim
,
Kim
,
2010
2004
Stock
, 2010
, 2010
, 1966 1
1
3
3
1
3
1
2
2
1
3
3
2
2
3
19
20
19
20
21
-
-
-
-
-
3
4
2
2
3
3
1
3
1
1
2
6
2
2
4
2
1
2
2
1
3
3
3
1
-
+
+ +
+
+ X
X
X
X
+
X
+ +
+ +
3= 1 1 3 1 2 3 3 3 1 1 1 X X, 3 - 3 18 17 18 18 - X X X X - Kim Kim Kim Kim Stock ((((2010 2010 2004 2010 (1966))) a))
A. quadridens Kim, 2016 1 1 3 20 - 3 3 4 2 1
+ X +
A
A
A
.
.
. rai reginae renaudi
Kim
Boxshal
Canu
& Min
, 1892
, 2013
& Huys, 1994 3
3
3 1
1
1 3
1
1
21
20
19 -
-
-
3
4
3
1
3
3
3
2
? 1
2
2
3
3
1 + + - X + - + + - 2 2 2 2 3 1 3 3 2 X - -, 3 18 17 - - X X - - Kim Kim Canu Boxshall (& 2016 (1892 Min &)) Huys (2013 () 1994)
A
A
A
.
.
. sarsi sensilis simplex
Bandera
Kim
Schirl
, 2010
&
, 1973
Conradi, 2009
3
4
1
1
1
1 3
2
4
20
21
20 -
-
- 3
3
4
2
1
3 4
2
2
1
2
2
2
3
-
X
+ -
X
X X + + - 2 2 2 3 3 3 X 2 - 17 17 - X X - Kim Schirl Bandera (2010 (1973 &) Conradi) (2014)
A
A
A
Neves
.
.
.
simulans siphonatus siphunculus
& Johnsson
(Scott
Giesbrecht
Bahia
T
, 2012
.,
,
1898
Canario
, 1897
)
, 4
1
2
1
1
3
3
2
4 21
20
20
-
- -
- 3
3
4
1
1
2 2
1
6 1
2
2 3
1
4
+ + + X X X X + + 1 2 1 3 3 3 X X - 18 18 - X X - Kim Bahia Present (2016 et study al). (2012)
A A
A.
..
spongus spinosus stimulans
Johnsson
Kim Giesbrecht & Min
, 2002
,, 1897 2013 2 4
3
2
3
1
3
3 4
18
20 21 -
-
3
3
4 1
3
-
3
5 3
1
1 2 1
1
3 + + + X X - + + - 1= - 2 2 3 3 - 1 X, - 2 17 18 - X + - Kim Johnsson Giesbrecht & Min (2002 (1899 (2013));) Bandera & Huys(2008)
A. stocki Nair & Pillai, 1984 1 1 2 20 - 2 1 6 1 1
+ X
A
A
A
A
.
..
.
tenerus suberitis tarifensis tenuicornis
(Hansen
Giesbrecht
Conradi
Brady
, 1923
,
&
1910
,
Bandera
1897
), 2011
4
5
3 3 3
2
2
1
-
3
2 2
21
21 21
21 -
-
-
-
4
4 4
4
? 3
2
1 3
2
2
2 5
2
2 2
2 3
1
3
3 + + + - X X X
X
X
+ +
+ +
1 2 1
1
1
3
3
3 3
3
1
2
-
-
- 17 18
18
-
-
X
X
X -
- Nair
Eiselt
Conradi Bandera Giesbrecht
&
(1965
Pillai
& & (Bandera Conradi
)
1899
(1984
); Bandera
)
((2011 2009) a) & Conradi (2009a)
A
A
A.
.
. tenuipes tricuspis tetrasetosus
Kim
Kim
Johnsson
,
,
2010
2010
, 1999
2
1
1
1
1
1 2
2
2
20
20
20 -
-
-
3
3
3 3
3
3 2
2
4
1
2
2
4
3
3
+ +
+
X
X X
X
+ + 1
2
3 3
3
2
1,
1 -
2
18
18
-
X
X - Kim
Kim
Johnsson
(
(2010
2010
(
1999)
)
)
A. trisetatus Kim, 2010 1 1 3 20 - 3 1 2 2 2 X
+ X 2 1 X 17 X Kim (2010)
A
A
A
A
A
.
.
..
.
walteri unioviger tubiporae urabensis uncinatus
Kim
(
Kim
Kim
Kim
Krichagin
, 2004
,
,
,
2010
2004
2004
, 1873)
3
1
1
3
4 1
1
1 1
1
2
2
3
3
3 19 21
21
19
21
-
-
-
- -
4
4
4
2
2 3
3
3
3
1
2
4 2
2
2
2
2
2 2
2
4 1
3
3
4 + +
+ + X X
X X
+
X
+ +
-
1=
2 1
2
3
2
3
3
2
3
1
1
X
, 3
1 -
3 18
17 18
18
- X X
+
- Kim Kim Kim Kim Marcus ((((2004 2004 2004 2010 & Por) a a b))) (1960, as A. carausi )
the genus Asterocheres are to be compared with new species described in this paper ( Table 2).
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